nomenclatural notes on tenebrionid beetles of the

Transcription

A N N A L E S Z O O L O G I C I (Warszawa), 2007, 57(3): 385-394
NOMENCLATURAL NOTES ON TENEBRIONID
BEETLES OF THE PALAEARCTIC REGION
(INSECTA: COLEOPTERA)
P. BOUCHARD1, I. LÖBL2 and O. MERKL3
1
Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture
and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON, K1A 0C6, Canada;
e-mail: [email protected]
2
Muséum d'histoire naturelle, Case Postale 6434, CH-1211, Geneva, Switzerland;
e-mail: [email protected]
3
Department of Zoology, Hungarian Natural History Museum, H-1088 Budapest,
Baross utca 13, Hungary; e-mail: [email protected]
Abstract.— This paper deals with nomenclatural problems in family-group and genusgroup names in Palaearctic Tenebrionidae. Triboliini, Amarygmini and Mycetocharina
were first made available in Faunula monacensis cantharologica (Gistel, 1848).
Neopachypterina nomen novum is proposed as a replacement name for Pachypterina
Medvedev, 1968 which is based on a junior homonym. The type genus of this subtribe is
Neopachypterus nomen novum (replacement name for Pachypterus Lucas, 1846, not
Swainson, 1839: Pisces). The available name Imatismus Dejean, 1834 is given priority over
Himatismus Erichson, 1843 and Curimosphena Gebien, 1920. A list of new combinations
for Imatismus is given. References are given to preserve the usage of Erodiini Bilberg,
1820, Epitragini Blanchard, 1845, Adesmiini Lacordaire, 1859, Metaclisa Jacquelin du Val,
1861 and Pachyscelis Solier, 1836 which are threatened by recently discovered older
names.
±
Key words.— Coleoptera, Tenebrionidae, darkling beetles, nomenclature, family-group
names, genus-group names.
INTRODUCTION
During the preparation of the Tenebrionidae section of the upcoming Catalogue of Palaeartic Coleoptera (Volume 5 – Tenebrionoidea), we have encountered nomenclatural problems in need of immediate
attention. Some of these require applications to the
International Commission on Zoological Nomenclature; these will be dealt with accordingly.
In this paper we first deal with nomenclatural notes
that supplement the recent treatment of family-group
names in Tenebrionidae (Bouchard et al. 2005). The
second part deals with three problems with genus-
group names that result in several new combinations in
one case and use of Article 23.9 (Reversal of Precedence) of the International Code of Zoological Nomenclature (ICZN 1999) to preserve a name in “prevailing
usage” in the others.
FAMILY-GROUP
NAMES
Change of author/year of publication
The following three family-group names published
in Gistel’s Faunula monacensis cantharologica
386
P. BOUCHARD, I. LÖBL and O. MERKL
predate names currently used as valid. The author
and/or date of publication need to be changed as follows.
Triboliini Gistel, 1848 (previously attributed to Mulsant, 1854).
Triboliidae Gistel, 1848a: [2]. Type genus: Tribolium MacLeay, 1825: 47.
Amarygmini Gistel, 1848 (previously attributed to
Gistel, 1856).
Amarygmiidae Gistel, 1848b: [2]. Type genus:
Amarygmus Dalman, 1823: 60.
Mycetocharina Gistel, 1848 (previously attributed to
Mulsant, 1856/Gistel, 1856).
Mycetocharisidae Gistel, 1848b: [2]. Type genus
Mycetochara Berthold, 1827: 371 (as Mycetochares Latreille, 1829).
Replacement names
The subtribe Pachypterina Medvedev, 1968, of the
tribe Opatrini Brullé, 1832, is permanently invalid and
therefore can no longer be used as valid because the
type genus Pachypterus Lucas, 1846 is a junior homonym. We hereby propose the following replacement
names:
Neopachypterina nomen novum. Comment: replacement name for Pachypterina Medvedev, 1968.
Type genus: Neopachypterus nomen novum
(replacement name for Pachypterus Lucas, 1846,
not Swainson, 1839: Pisces).
Preservation of names in prevailing usage
Three family-group names currently used as valid
are threatened by recently discovered older names.
Because the older names were not used as valid after
1899 to our knowledge, we present supporting references to preserve the currently accepted names following the requirements of Article 23.9 (see Appendices
1–3).
Erodiini Billberg, 1820 nomen protectum
Erodiides Billberg, 1820: 32. Comment: name made
available under Art. 11.7.2. Type genus: Erodius
Fabricius, 1775: 258.
Cephacerini Rafinesque, 1815 nomen oblitum
Cephaceria Rafinesque, 1815: 113. Type genus:
Cephacerus Rafinesque, 1815: 113 (unjustified
emendation of Erodius Fabricius, 1775).
Epitragini Blanchard, 1845 nomen protectum
Épitragites Blanchard, 1845: 16. Comment: name
made available under Art. 11.7.2. Type genus:
Epitragus Latreille, 1802: 165.
Lygophilini Rafinesque, 1815 nomen oblitum
Lygophilia Rafinesque, 1815: 113. Type genus:
Lygophilus Rafinesque, 1815: 114 (unjustified
emendation of Epitragus Latreille, 1802).
Adesmiini Lacordaire, 1859 nomen protectum
Adesmiides Lacordaire, 1859: 22. Comment: name
Adesmia Fischer von Waldheim, 1822: 153.
Megageniini Solier, 1851 nomen oblitum
Megagenianos Solier, 1851: 124. Comment: name
Megagenius Solier, 1835: 513.
GENUS-GROUP
NAMES
Imatismus Dejean – Himatismus Erichson –
Curimosphena Gebien
Imatismus was established by Dejean, 1834: 202
who included only one valid species, [Helops] fasciculatus Fabricius, 1798. This species is the type of the
genus by monotypy. Curimosphena Gebien, 1920 is
currently considered a valid subgenus of Himatismus
Erichson, 1843, and includes Helops fasciculatus
Fabricius, 1798. The following nomenclatorial changes
result from these facts: Curimosphena Gebien is
a junior synonym with Imatismus Dejean; Himatismus Erichson is downgraded to subgenus of Imatismus Dejean, and the taxa previously placed in
Himatismus Erichson, 1843 are transferred to Imatismus Dejean. Imatismus includes the following
valid species-group taxa:
genus Imatismus Dejean, 1834: 202 type species
Helops fasciculatus Fabricius, 1798
subgenus Imatismus Dejean, 1834: 202 type species
Helops fasciculatus Fabricius, 1798
=Curimosphena Gebien, 1920: 43 type species Himatismus villosus Haag-Rutenberg, 1870, syn. nov.
abyssinicus (Koch, 1950) comb. nov. from Himatismus
angolanus (Koch, 1950) comb. nov. from Himatismus
arabicus (Kaszab, 1981) comb. nov. from Himatismus
barnardi (Koch, 1950) comb. nov. from Himatismus
basilewskyi (Koch, 1950) comb. nov. from Himatismus
baudii (Haag-Rutenberg, 1877) comb. nov. from
Himatismus
NOMENCLATURE OF PALAEARCTIC TENEBRIONIDAE
brevicornis (Koch, 1962) comb. nov. from Himatismus
buprestoides buprestoides (Gerstaecker, 1854)
comb. nov. from Himatismus
buprestoides cylindricollis (Koch, 1950) comb. nov.
from Himatismus
buprestoides strangulatus (Koch, 1950) comb. nov.
from Himatismus
chinensis (Reitter, 1916) comb. nov. from Himatismus
congolanus (Koch, 1950) comb. nov. from Himatismus
dahomeyus (Koch, 1950) comb. nov. from Himatismus
dubia (Haag-Rutenberg, 1870) comb. nov. from
Himatismus
epitragoides (Haag-Rutenberg, 1870) comb. nov.
from Himatismus
eclavata (Lesne, 1920) comb. nov. from Himatismus
fasciculatus (Fabricius, 1798)
ferreirai (Koch, 1952) comb. nov. from Himatismus
gedyei (Koch, 1950) comb. nov. from Himatismus
gomesi (Koch, 1950) comb. nov. from Himatismus
haroldi (Haag-Rutenberg, 1878) comb. nov. from
Himatismus
heterophthalmus heterophthalmus (Chatanay,
1914) comb. nov. from Himatismus
heterophthalmus transbechuanus (Koch, 1950)
incisa (Gebien, 1937) comb. nov. from Curimosphena
inconspecta (Haag-Rutenberg, 1877) comb. nov. from
Himatismus
justi (Fairmaire, 1880) comb. nov. from Himatismus
lindneri (Kolbe, 1883) comb. nov. from Himatismus
luctuosus (Fairmaire, 1895) comb. nov. from Himatismus
ludwigi (Carl, 1993) comb. nov. from Himatismus
maculisparsa (Lesne, 1920) comb. nov. from Himatismus
marginatus (Koch, 1950) comb. nov. from Himatismus
muelleri (Haag-Rutenberg, 1877) comb. nov. from
Himatismus
natalensis natalensis (Baudi, 1875) comb. nov. from
Himatismus
natalensis pinheyi (Koch, 1850) comb. nov. from
Himatismus
nigrosulcata (Quedenfeld, 1885) comb. nov. from
Himatismus
occidentalis (Lacordaire, 1876) comb. nov. from
Himatismus
ovipennis (Koch, 1950) comb. nov. from Himatismus
387
patruelis capensis (Koch, 1950) comb. nov. from
Himatismus
patruelis patruelis (Bertolini, 1849) comb. nov. from
Himatismus
pavesi (Gestro, 1892) comb. nov. from Himatismus
planicollis (Kolbe, 1883) comb. nov. from Himatismus
plebeja (Fåhraeus, 1870) comb. nov. from Himatismus
recticollis (Koch, 1950) comb. nov. from Himatismus
rileyi (Koch, 1953) comb. nov. from Himatismus
rhodesianus (Koch, 1950) comb. nov. from Himatismus
schoutedeni (Koch, 1950) comb. nov. from Himatismus
scorteccii (Koch, 1962) comb. nov. from Himatismus
semialatus (Koch, 1950) comb. nov. from Himatismus
seriata (Gebien, 1937) comb. nov. from Curimosphena
senegalensis ennediansis (Español, 1973) comb.
nov. from Himatismus
senegalensis senegalensis (Haag-Rutenberg, 1870)
somalicus (Koch, 1950) comb. nov. from Himatismus
siffointei (Ardoin, 1974) comb. nov. from Himatismus
sublinearis (Walker, 1871) comb. nov. from Himatismus
terryi coniops (Koch, 1950) comb. nov. from Himatismus
terryi swazinus (Koch, 1950) comb. nov. from
Himatismus
terryi terryi (Koch, 1950) comb. nov. from Himatismus
tessulatus (Gerstæcker, 1854) comb. nov. from
Himatismus
transvaalensis rotundophthalmus (Koch, 1950)
transvaalensis transvaalensis (Koch, 1950) comb.
nov. from Himatismus
trivialis septentrionalis (Lesne, 1922) comb. nov.
from Himatismus
trivialis trivialis (Gerstæcker, 1871) comb. nov.
from Himatismus
trochanterica (Fairmaire, 1897) comb. nov. from
Himatismus
vaneyeni (Ardoin, 1974) comb. nov. from Himatismus
vansoni karrooensis (Koch, 1950) comb. nov. from
Himatismus
vansoni vansoni (Koch, 1950) comb. nov. from
Himatismus
388
variegata karrooensis (Koch, 1950) comb. nov. from
Himatismus
variegata vansoni (Koch, 1950) comb. nov. from
Himatismus
variegata (Fabricus, 1781) comb. nov. from Himatismus
venzoi (Koch, 1962) comb. nov. from Himatismus
vestita Baudi di Selve, 1875) comb. nov. from Himatismus
villosus (Haag-Rutenberg, 1870) comb. nov. from
Himatismus
whiteheadi (Ferrer, 1995) comb. nov. from Himatismus
therefore present supporting references to preserve
the younger genus name Pachyscelis Solier, 1836
which is currently used as valid (Appendix 5).
subgenus Himatismus Erichson, 1843: 255 type species Himatismus mandibularis Erichson, 1843
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present supporting references to preserve the younger
genus name Metaclisa Jacquelin du Val, 1861 which is
currently used as valid (Appendix 4).
Brachyscelis Dejean – Pachyscelis Solier
Dejean (1834: 179) made the genus name Brachyscelis available by including three available species
(Pimelia metopotapha Ménétriés, 1832, P. clavaria
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placed in Pachyscelis Solier, 1836. The older name
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well-established name Pachyscelis Solier. In addition,
the genus name Brachyscelis Germar, 1834 is currently used as valid in the family Chrysomelidae, giving even more reason to suppress Dejean’s name.
The name Brachyscelis Dejean, 1834 has not been
used as valid in the literature to our knowledge and we
ACKNOWLEDGMENTS
We thank M. Schmitt (Zoologisches Forschungsmuseum Alexander Koenig) for his comments on the
usage of Brachyscelis Germar in Chrysomelidae. We
also thank Y. Bousquet and A. Smetana (Canadian National Collection of Insects) for reviewing this manuscript.
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Tenebrionidae). The Coleopterists’ Bulletin, 55: 515–521.
Doyen, J. T. 1974. Larvae of Bothrotes plumbeus and Lobometopon fusiforme (Coleoptera: Tentyriidae: Epitragini)
The Coleopterists Bulletin, 28: 159–165.
Doyen, J. T. and W. R. Tschinkel. 1982. Phenetic and cladistic
Doyen, J. T. 1989. Tenebrionidae and Zopheridae of the Chamela Biological Station and vicinity, Jalisco, Mexico
(Coleoptera). Folia Entomologica Mexicana, 77 (1988):
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Doyen, J. T. 1994. Cladistic relationships among Pimeliine
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Ferrer, J. 2001. Une nouvelle espèce d’Aphrotus Péringuey,
1899 du Botswana (Coleoptera, Tenebrionidae, Epitragini). Entomofauna, 22: 9–14.
Ferrer, J. 2005. Sur la vraie position du genre Leprocaulinus
Kaszab, 1982. (Coleoptera, Tenebrionidae, Epitiragini).
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Ferrer, J. and J. H. Yvinec. 2004. Révision de la tribu des
Lachnogyini Reitter, 1904 sensu nov. et description d’un
nouveau genre et d’une espèce nouvelle du désert de Taklamakan, Chine (Coleoptera: Tenebrionidae, Pimeliinae).
Annales de la Société Entomologique de France, 40: 41–49.
Freude, H. 1968. Revision der Epitragini (Coleoptera-Tenebrionidae). Entomologische Arbeiten aus dem Museum Georg
Frey, 19: 32–143.
Freude, H. 1981. Berichte über eine Determinasionssendung
von Herrn Luis E. Peña G. mit Neubeschreibungen von
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Freude, H. 1986. Drei neue Epitragini-Arten aus Guatemala
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Freude, H. 1993. Neue Monommidae und Epitragini (Tenebrionidae) des British Museum und eine Tabelle der
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Marcuzzi, G. 1983. Description of the external male genitalia
of some Neotropical Tenebrionidae (Col. Heteromera).
Folia Entomologica Hungarica, 44: 239–269.
Marcuzzi, G. 1984. A catalogue of tenebrionid beetles (Coleoptera: Heteromera) of the West Indies. Folia Entomologica
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Marcuzzi, G. 1987. Description of the external female genitalia
of some Neotropical Tenebrionidae (Coleoptera: Heteromera). Acta Zoologica Academiae Scientiarum Hungaricae, 33: 87–112.
Merkl, O. 1991. Epitragini from Iran (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 37: 59–66.
Peck, S. B. 2005. A checklist of the beetles of Cuba with data
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Gainesville. vi + 241pp.
Ren, G. and Z. Zheng. 1993. Four mew [sic] species of tribe
Epitragini from desert in northwest of China (Coleoptera:
Tenebrionidae). Journal of Ningxia Agricultural College,
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392
Appendix 3.
Supporting references for the preservation of Adesmiini
Lacordaire, 1859 over Megageniini Solier, 1851 under Article
23.9. The tribe name Megageniini Solier, 1851 has not been
used as valid after 1899 to our knowledge.
Alfieri, A. 1976. The Coleoptera of Egypt. Mémoires de la
Société Entomologique d’Égypte, 5: 1–361.
Cloudsley Thompson, J. L. 2001. Thermal and water relations
of desert beetles. Naturwissenschaften, 88: 447–460.
Doyen, J. T. 1994. Cladistic relationships among Pimeliine
Tenebrionidae (Coleoptera). Journal of the New York
Entomological Society, 101 (1993): 443–514.
Duncan, F. D. 2003. The role of the subelytral cavity in respiration in a tenebrionid beetle, Onymacris multistriata
(Tenebrionidae: Adesmiini). Journal of Insect Physiology,
49: 339–346.
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Ferreira, M. C. 1965. Catálogo dos Coleópteros de Angola.
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Hanrahan, S. A. and S. W. Nicolson. 1984. The effect of dehydration on the Malpighian tubules of Onymacris plana
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(Coleoptera: Tenebrionidae: Adesmiini) in the central
Namib Desert dunes. Journal of the Entomological Society
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Kaszab, Z. 1981. Insects of Saudi Arabia. Coleoptera: fam.
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Koch, C. 1962. The Tenebrionidae of Southern Africa. XXXI.
Comprehensive notes on the tenebrionid fauna of the
Namib desert. Annals of the Transvaal Museum, 24:
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Kwieton, E. 1982. Revue critique des systèmes récents de la
famille des Tenebrionidae (Col.). Acta Musei Nationalis
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McClain, E., Praetorius, R. L., Hanrahan, S. A. and M. K. Seely.
1984. Dynamics of the wax bloom of a seasonal Namib
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Penrith, M.-L. 1979. Revision of the western southern African
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Series A, 5: 1–94.
Penrith, M.-L. 1986. Relationships in the tribe Adesmiini
(Coleoptera: Tenebrionidae) and a revision of the genus
Stenodesia Reitter. Annals of the Transvaal Museum, 34:
275–302.
Roer, H. 1981. Weitere Untersuchungen zur Anpassung des
Namibwüstenkäfers Onymacris r. rugatipennis (Haag
1875, Col.: Tenebrionidae, Adesmiini) am das Trockenflussbett des Kuiseb in Südwestafrika. Mitteilungen der
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Roer, H. 1983. Aktionsraum und Anpassungsphanomene des
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Roer, H. 1985. Dispersion dynamics of tenebrionids of the
genus Onymacris (Col.: Tenebrionidae, Adesmiini) in the
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Roer, H. 1986. Zur Anpassung des Schwarzkafers Onymacris unguicularis (Haag) (Col.: Tenebrionidae, Adesmiini) an die Nebelzone der Namibwüste. Bonner Zoologische
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Skopin, N. G. 1978. Tenebrionidae, pp. 223–266. In: Klausnitzer, B. (ed.), Ordnung Coleoptera (larven). W. Junk,
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Veiga Ferreira, G., da. 1964. Catálogo dos tipos de insectos
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393
Appendix 4.
Supporting references for the preservation of Metaclisa
Jacquelin du Val, 1861 over Amarantha Motschulsky, 1859
under Article 23.9. The genus name Amarantha Motschulsky, 1859 has not been used as valid after 1899 to our knowledge.
Aalbu, R. L., Flores, G. E. and C. A. Triplehorn. 2002. Tenebrionidae, pp. 499–512. In: Bousquets, J. L. and Morrone, J.
J. (eds.), Biodiversidad, taxonomia y biogeografia de
arthropodos de Mexico: Hacia una sintesis de su conocimiento. Vol. III. Universidad Nacional Autonoma de Mexico, Mexico.
Aalbu, R. L., Triplehorn, C. A., Campbell, J. M., Brown, K. W.,
Somerby, R. E. and D. B. Thomas. 2002. 106. Tenebrionidae Latreille 1802, pp. 463–509. In: Arnett, R. H. Jr.,
Thomas, M. C., Skelley, P. E. and J. H. Frank (eds.), American Beetles. CRC Press, New York.
Arnett, R. H. Jr. 1963. The beetles of the United States (a manual for identification). The Catholic University of America
Press, Washington D.C. 1112 pp.
Boddy, D. W. 1965. Family Tenebrionidae, pp. 130–189. In:
Hatch, M. H. (ed.), The beetles of the Pacific northwest.
Part IV: Macrodactyles, Palpicornes, and Heteromera.
University of Washington Press, Seattle.
Bousquet, Y. and J. M. Campbell. 1990. Family Tenebrionidae
darkling beetles, pp. 253–261. In: Bousquet, Y. (ed.),
Checklist of the beetles of Canada and Alaska. Research
Branch, Ottawa.
Chûjô, M. 1968. A check list of the formosan Tenebrionidae
(Coleoptera) III. Kontyû, 36: 393–397.
Chûjô, M. 1977. Tenebrionidae of the Nansei Islands II (Coleoptera). Esakia, 10: 1–19.
Chûjô, M. 1978. Tenebrionidae of the Nansei Islands III (Coleoptera). Esakia, 11: 63–80.
Chûjô, M. 1978. Tenebrionidae of the Nansei Islands IV (Coleoptera). Esakia, 12: 1–16.
Chûjô, M. 1979. Tenebrionidae of the Nansei Islands V (Coleoptera). Esakia, 14: 31–56.
Doyen, J. T. 1984. Reconstitution of the Diaperini of North
America, with new species of Adelina and Sitophagus
(Coleoptera: Tenebrionidae). Proceedings of the Entomological Society of Washington, 86: 777–789.
Doyen, J. T. 1989. Reconstitution of Coelometopini, Tenebrionini and related tribes of America north of Colombia (Coleoptera: Tenebrionidae). Journal of the New York Entomological Society, 97: 277–304.
Doyen, J. T. and Tschinkel, W. R. 1982. Phenetic and cladistic
Hirashima, Y. 1989. A check list of Japanese insects. Volume
I. Kyushug Daigaku Nogakubu Konchu Kyoshitsu, Fukuoka. xiv + 540 pp.
Kim, S. Y. and J. I. Kim. 2002. Newly recorded and little known
tenebrionid species from Korea. Korean Journal of Entomology, 32: 243–246.
Kompantseva, T. V. 1997. The larva of Metaclisa viridis (Motschulsky, 1859) and notes on the position of the genus
Metaclisa Jacquelin du Val, 1861 in the family Tenebrionidae (Coleoptera). Entomologicheskoe Obozrenie, 76:
606–612. [In Russian: English translation in Entomological
Review, 76(3): 636–642].
Li-zhong, H. 2002. List of Chinese insects. Volume II. Zhongshan (Sun Yat-sen) University Press, Guangzhou. 612 pp.
Nakane, T. 1963. Tenebrionidae, pp. 220, pls 111–118. In:
Nakane, T., Ohbayashi, K., Nomura, S. and Y. Kurosawa.
(eds.), Iconographia Insectorum Japonicum Colore naturali edita. Volumen II (Coleoptera). Hokuryukan, Tokyo.
Nakane, T. 1968. New or little-known Coleoptera from Japan
and its adjacent regions. XXVII. Fragmenta Coleopterologica Japanica, 19: 76–78; 20: 79–82; 21: 83–85.
Shibata, T. 1978. Notes on the Tenebrionidae from Taiwan
and Japan, I. (Coleoptera). Entomological Review of Japan, 32: 19–27.
Triplehorn, C. A. 1965. Revision of Diaperini of America north
of Mexico with notes on extralimital species (Coleoptera:
Tenebrionidae). Proceedings of the United States National Museum, 117: 349–458.
Tschinkel, W. R. 1975a. A comparative study of the chemical
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secretions. Journal of Insect Physiology, 21: 753–783.
Tschinkel, W. R. 1975b. A comparative study of the chemical
defensive system of Tenebrionid beetles. Defensive behaviour and ancillary features. Annals of the Entomological
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Tschinkel, W. R. 1975c. A comparative study of the chemical
defensive system of tenebrionid beetles. III. Morphology of
the glands. Journal of Morphology, 145: 355–370.
Tschinkel, W. R. and Doyen, J. T. 1980. Comparative anatomy of the defensive glands, ovipositors and female
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394
Appendix 5.
Supporting references for the preservation of Pachyscelis Solier, 1836 over Brachyscelis Dejean, 1834 under
Article 23.9. The genus name Brachyscelis Dejean, 1834 has
not been used as valid after 1899 to our knowledge.
Allsopp, P. G. 1980. The biology of false wireworms and their
adults (soil-inhabiting Tenebrionidae) (Coleoptera): a review. Bulletin of Entomological Research, 70: 343–379.
Bogachev, A. V. and M. G. Nepesova. 1980. Tenebrionid beetles
of the genus Pachyscelis Sol. (Coleoptera, Tenebrionidae) from Turkmenia. Entomologicheskoe Obozrenie, 59:
607–612. [In Russian: English translation in Entomological
Review, 59(3): 84–89].
Dajoz, R. 1976. Notes écologiques et biogéographiques sur
quelques coléoptères Tenebrionidae de Grèce. L’Entomologiste, 32: 174–179.
Holecova, M., Lachowska, D. and G. Karagyan. 2002. Karyological notes on six beetle species from Armenia (Coleoptera: Tenebrionidae, Cerambycidae, Curculionidae).
Folia Biologica, 50: 9–12.
Kaszab, Z. 1958. Ergänzungen zur Tenebrioniden (Insecta).
Aufarbeitung von Dr. E. Goidelli der 3. danischen zentralasiatischen Expedition. (Zool. Results 24). Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening,
Copenhagen, 120: 237–255.
Kaszab, Z. 1960. Die Tenebrioniden Afghanistans auf Grund
der Ergebnisse des Sammelreise des Herrn J. Klapperich
in den Jahren 1952–53 (Col.) (End.). Entomologische
Arbeiten aus dem Museum Georg Frey, 11: 1–179.
Kaszab, Z. 1969. Beiträge zur Kenntnis der Fauna
Afghanistans. (Sammelergebnisse von O. Jakeš, 1963–64,
D. Povolný, 1965, D. Povolný & Fr. Tenora, 1966, J. Šimek,
1965–66, D. Povolný, J. Gaisler, Z. Šebek & Fr. Tenora,
1967) Tenebrionidae, Col. Acta Musei Moraviae, 54: 2–182.
Kaszab, Z. 1973. Neue Beiträge zur Kenntnis der Tenebrioniden-Fauna von Afghanistan (Coleoptera). Entomologica
Scandinavica, 4: 35–58.
Kwieton, E. 1977. Esquisse phylogenetique du genre Pimelia
F. Acta Entomologica Musei Nationalis Pragae, 39:
559–589.
Kwieton, E. 1978. Sur la division de l’ancien genre Pachyselis
Sol. (Col., Tenebrionidae). Bulletin de la Société Entomologique de Mulhouse, 34: 28–34.
Kwieton, E. 1980. Révision du genre Pachyscelis Solier et
notes sur des relations inter-génériques chez les Pimellini
(Col., Tenebrionidae). Annotationes Zoologicae et Botanicae, 137: 1–16.
Kwieton, E. 1987. Mise au point sur la phylogenie et la paleogeographie du g. Pimelia F. (Col. Tenebrionidae). Bulletin de la Société Entomologique de Mulhouse. Supplément, 15: 9–15.
Löbl, I. and O. Merkl. 2003. On the type species of several tene-
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Medvedev, G. S. 1964. New forms of darkling beetles (Coleoptera, Tenebrionidae) from soviet Central Asia. Entomologicheskoe Obozrenie, 43: 651–661. [In Russian: English
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Medvedev, G. S. 1992. A new species of the genus Pachyscelis
Sol. (Coleoptera, Tenebrionidae) from southern Kyrgyzstan. Entomologicheskoe Obozrenie, 71: 793–795, 951. [In
Russian: English translation in Entomological Review, 72:
79–80].
Medvedev, G. S. and M. G. Nepesova. 1985. Key to the identification of the darkling beetles of Turkmenistan. Ylym,
Ashkhabad. 178 pp. [In Russian].
Pierre, F. 1965. Contribution à la connaissance de la faune du
Moyen-Orient (Missions G. Remaudiere 1955 et 1959) II.
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Pierre, F. 1968. Contribution à la faune de l’Iran. 12 Étude
écologique et biogéographique des Ténébrionides. Pimeliinae s. nov. des régions désertiques et semi-désertiques de
l’Iran. Annales de la Société Entomologique de France, 4:
997–1036.
Pierre, F. 1981. A propos de la distribution et de la variation
des espèces dans le genre Pachyscelis (s.l.). (Col. Tenebrionidae). Bulletin de la Société Entomologique de
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Slilvano, C. 1968. Su qualche Pachyscelis di Persia. (18 Contribute allo studio dei tenebrionidi). Bollettino della Società Entomologica Italiana, 98: 133–135.
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Tezcan, S., Ferrer, J. and B. Keskin. 2000. Contribution to the
study of tenebrionid beetles (Coleoptera: Tenebrionidae)
in ecological cherry orchards in Ýzmir and Manisa
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Winkler, J. R. and I. Zapletalová. 1974. Some remarks on Pachyscelis zhenzhuristi Bogatschev, 1939 (Coleoptera:
Tenebrionidae), endemic species of Afghanistan: introduced to Czechoslovakia with imported raisins. Zborník
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Yaacobi, G., Ziv, Y. and M. L. Rosenzweig. 2007. Effects of
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nomenclatural notes on tenebrionid beetles of the

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