The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae

Transcription

The Madagascan Coraebina Bedel, 1921 (Coleoptera:
Buprestidae: Agrilinae: Agrilini): introduction and
systematic catalogue*
C. L. Bellamy†
Transvaal Museum, P.O. Box 413, Pretoria, 0001 South Africa
Bellamy, C. L., 2001. The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae: Agrilinae: Agrilini): introduction and systematic catalogue. Annals of the Transvaal Museum 38: 57–70.
An introduction to a vast descriptive and systematic project that will consider the large fauna of coraebine jewel
beetles from Madagascar and the Comoro Islands is presented. The described species are summarized in a
catalogue that contains 28 genera, one subgenus, 103 species and one subspecies. Coraebastus imperatrix
(Obenberger), from Madagascar, is resurrected from synonymy under C. quinquepustulatus Fairmaire, from the
Comoro Islands. Chloricala gratiosa Kerremans var. octosticta Obenberger and var. impunctata Théry and
Pseudagrilus scintillans var. madegassus Obenberger are synonymized under their nominate species.
Le vaste projet de description systématique de l’importante faune des Buprestes Coraebinae de Madagascar et des
Comores est présenté. La description des espèces est résumée dans un catalogue contenant 28 genres, un
sous-genre, 103 espèces et une sous-espèce. Coraebastus imperatrix (Obenberger), de Madagascar, est rétabli au
rang d’espèce. Il n’est plus synonyme de C. quinquepustulatus Fairmaire des Comores. Les var. otosticta
Obenberger et impunctata Théry du Chloricala gratiosa Kerremans ainsi que le Pseudagrillus scintillans var.
madegassus Obenberger sont synonymisés sous le nom de leurs espèces nominales.
Keywords/Mots Clés: Coleoptera, Buprestidae, Agrilinae, Coraebina, Madagascar.
INTRODUCTION
There are many accounts of a proliferation of
species in the literature. Perhaps these come more
frequently now with more attention being paid to
inventorying threatened habitats as world populations rise and development invades once pristine
tracts of nature. In my experience, there is nothing
that compares to the vast and largely undescribed
jewel beetle subtribe Coraebina Bedel (1921)
(Coleoptera: Buprestidae: Agrilinae: Agrilini) from
Madagascar. This is the second contribution in a
series that is anticipated to take many years to
complete; the first was by Bellamy (1997).
During the latter half of the 20th century, French
and Malagasy entomologists discovered that
nest-provisioning wasps (Hymenoptera: Sphecoidea: Philanthidae: Cerceris Latreille) were using
small jewel beetles as their primary prey. The two
principal collectors on the majority of specimen
data labels are P. Vadon and A. Peyrieras. The beetles are paralyzed by the sting of female wasps and
then carried back to the nests, whereupon an egg is
laid and the emerging larva feeds on the beetle
before pupating inside the hollow body. By following
the wasps and learning to recognize their burrows
*Third contribution to the knowledge of Madagascar Buprestidae.
†
Present address: Entomology Section, Natural History Museum of
Los Angeles County, 900 Exposition Blvd., Los Angeles, California,
90007, U.S.A. E-mail: [email protected]
or nests, these entomologists were able to dig up
many comatose beetles, which lay in perfect or
near-perfect condition, and return them for preparation and permanent museum storage.
The predator–prey relationship between certain
species of Cerceris and buprestids has been widely
reported for various parts of the world (e.g. in general: Cartwright, 1931; Inglebert, 1993; Scullen and
Wold, 1969; Trinidad: Fisher, 1944; Callan, 1990;
North America: Grossbeck, 1912; Linsley and
MacSwain, 1956; Evans and Rubinik, 1978;
Kurczewski and Miller, 1984; Hook and Evans,
1991; Japan: Ohmomo and Usuba, 1997).
Several colleagues (W. F. Barr, H. A. Hespenheide,
S. G. Wellso, pers. comm.) had told me about the
collection of Madagascan buprestids seen in the
Museum National d’Histoire Naturelle, Paris, during
their respective research visits in the 1970s and
1980s, but until I experienced this firsthand during
my initial visit to the museum in late 1995, I really
could not understand the breadth of this collection.
There are thousands of specimens, prepared and
labelled, representing a large number of mostly
undescribed species. From an initial attempt to
sort and arrange these specimens by the former head of the Coleoptera Department, the late
A. Descarpentries, an estimate of the size of this
fauna is between 500–600 undescribed species,
which may require 20–30 new genera to contain
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ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001
them. It seems clear from the literature, as well as
from the examples of other buprestid prey in this
material, that the Cerceris predators are not
particular to coraebine buprestids, nor possibly
even just buprestids. There are large series of other
buprestid species (e.g., Sponsor Gory and Laporte,
see Descarpentries, 1965; Anthaxia Eschscholtz,
see Descarpentries, 1967) that are each of about
the same general size and shape as the many
coraebine species and this suggests that the
predators are perhaps more selective about the
physical characteristics of the prey, perhaps to be
able to contain them within specific morphological
adaptations to maintain a hold. Krombein (1981: 8)
correlated structure and function when he described and figured the ‘buprestid clamp’ found in
members of the ‘bupresticida group’ of Cerceris,
including species from Sri Lanka, Europe and North
America.
The present paper is intended as an introduction
to the many nomenclatural acts and descriptive
papers that must follow. By introducing the subject
and providing an accounting of the current systematics of the known Madagascan coraebine fauna, all
subsequent efforts can focus on the description of
new taxa and several of the large genera which have
never been revised, e.g. Entomogaster Saunders.
The catalogue included herein is presented in full,
indicating all taxa and the nomenclatural history for
each species. This effort to present a full catalogue
of the previously described taxa of this subtribe is
undertaken now to bring together all of the available
information on a large group of species that have
mostly appeared in random and disparate descriptive publications and earlier more general catalogues. The chronology of catalogues that have
treated these taxa, in part, are: Gemminger and Harold, 1869; Saunders, 1871; Kerremans, 1885, 1892;
Alluaud, 1900; Kerremans, 1903; Théry, 1905; Marie
and Lesne, 1917; Obenberger, 1934; and Bellamy,
1985. The work of Obenberger (1934) was the fourth
of six fascicles of the two volumes on Buprestidae in
the Coleopterorum Catalogue, the last effort to catalogue the world’s beetle fauna. Many of the taxa of
Madagascan Coraebina were described subsequent to that catalogue and have not been found in
the literature after their respective original descriptions.
For future geographical notes, distributional
accounts and attempts to summarize the range of
particular taxa within the Madagascan habitats, I
note here that a great many of the older taxa of this
group were described from a type locality no more
precise than ‘Madagascar’. However, considering
the wealth of specimens of many of these taxa that
were harvested, some detail will be filled in as
species are described and genera are revised. As a
reference for Madagascan place names, the work
of Viette (1991) will be the standard.
The described complement of these beetles is 28
genera, one subgenus, 103 species and one subspecies. The single related taxon, Coraebastus
quinquepustulatus Fairmaire, from the Comoro
Islands is included with the reinstatement from
synonymy of Coraebastus imperatrix (Obenberger)
from Madagascar. In the catalogue, the genera and
species are listed alphabetically. With so much
descriptive work to be done, phylogenetic predictions are premature, since each subsequent
descriptive paper will alter the conclusions about
transformation character state series and
synapomorphies. At the end of the catalogue are
listed 11 species names that were originally
described in one of two older coraebine genera.
These are listed as ‘undefined placement’ pending
better understanding of relationships with the many
new species amongst which some of these will
undoubtedly fit. Following current trends in dealing
with infrasubspecific names, I will propose all
varietal names listed below as new synonyms (see
Bellamy & Peterson, 2000).
Abbreviations used herein are: nom. nov. =
nomen novum; nom. nud. = nomen nudum; nom.
superfl. = superfluous name, i.e. a replacement
name that was unnecessary; orig. desc. = original
description; rev. = revision; spp. key = species key;
syn. = synonym; syn. nov. = new synonym;
unvn. = unavailable name, i.e. infrasubspecific
names, nomina nuda; var. = variety, variation.
SYSTEMATIC CATALOGUE OF THE
COROEBINA OF THE MADAGASCAN
REGION
Genus 1. ANGATRA Descarpentries
Angatra Descarpentries, 1969: 599–600 (orig.
desc.); Bellamy, 1985: 424.
Type species: Angatra magna Descarpentries
(original designation).
REMARKS. This genus is endemic to Madagascar
and apparently has no close relationships to the
Afrotropical fauna.
1. magna Descarpentries
Angatra magna Descarpentries, 1969: 600–601
(orig. desc.), fig. 1.
2. parva Descarpentries
Angatra parva Descarpentries, 1969: 601–602
Genus 2. ANTANAMBIA Descarpentries
Antanambia Descarpentries, 1975: 19–20 (orig.
MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE
Ty p e s p e c ie s : A n t a n a m b i a n i g r i c a ns
Descarpentries (original designation).
and it is not particularly close to any coraebine
lineage in sub-Saharan Africa.
3. nigricans Descarpentries
Antanambia nigricans Descarpentries, 1975:
21–22 (orig. desc.), figs 1, 3–5.
Genus 3. BRACHYDORA Obenberger
Brachydora Obenberger, 1923: 30–31 (orig.
desc.); 1934: 794; Bellamy, 1985: 423.
Ty p e s p e c ie s : B r a c h y d or a m o ns t r um
Obenberger (original designation).
and is quite distinct from the Afrotropical fauna.
4. crassa Théry
Brachydora crassa Théry, 1937: 87–89 (orig.
desc.), fig. 3.
5. deformis (Fairmaire)
Discoderes deformis Fairmaire, 1901: 163 (orig.
desc.); Kerremans, 1903: 223; Marie and
Lesne, 1917: 40.
Paradora deformis: Théry, 1905: 125.
Brachydora deformis: Obenberger, 1923: 32;
1934: 795.
6. granulum (Fairmaire)
Discoderes granulum Fairmaire, 1902b:
350–351 (orig. desc.); Théry, 1905: 116;
Marie and Lesne, 1917: 40
Brachydora granulum: Obenberger, 1928: 342;
1934: 795.
7. monstrum Obenberger
Brachydora monstrum Obenberger, 1923: 31–32
(orig. desc.); 1934: 795.
8. sicardi (Théry)
Paradora sicardi Théry, 1912: 118–119 (orig.
desc.,)
Brachydora sicardi: Obenberger, 1923: 31; 1934:
795.
Genus 4. CHLORICALA Kerremans
Chloricala Kerremans, 1893: 119–120 (orig.
desc.); Théry, 1905: 120; Bellamy, 1985: 424.
Type species: Chloricala gratiosa Kerremans
(original monotypy).
Chloricalla (lapsus calami): Kerremans, 1900:
319; 1903: 231.
59
and shows no close similarities to the Afrotropical
fauna. The two varietal names deserve no recognition and are technically invalid (ICZN 1999, Article
45.5, 45.6).
9. balachowskyi Descarpentries
Chloricala balachowskyi Descarpentries, 1974:
582–584 (orig. desc.), figs 7, 8.
10. gratiosa Kerremans
Chloricala gratiosa Kerremans, 1893: 120 (orig.
desc.); 1900: 319; 1903: 231, plate 2, fig. 11;
Théry, 1905: 120; Obenberger, 1934: 813;
1935b: 20; Théry, 1936a: 59.
Chloricala gratiosa var. octosticta Obenberger,
1935b: 20 (orig. desc.); 1934: 813; Théry,
1936a: 59. Syn. nov.
Chloricala gratiosa var. impunctata Théry, 1936a:
60–61 (orig. desc.). Syn. nov.
Genus 5. COMPSOGLYPTA Fairmaire
Compsoglypta Fairmaire, 1904: 233 (orig.
desc.); Kerremans, 1910: 3 (syn. Psiloptera),
347; Marie and Lesne, 1917: 30; Obenberger,
1928: 339; 1934: 795; Bellamy, 1985: 423.
Type species: Compsoglypta perrieri Fairmaire
Compsoglypha (lapsus calami): Théry, 1905:
177; Obenberger, 1934: 75; Bellamy, 1985:
423.
and its placement in Psilopterina or Coraebina has
been somewhat confused as indicated by the
synonymy above. I have examined the types of the
Fairmaire taxon and am confident that this is a
member of the coraebine lineage, albeit very divergent.
11. perrieri Fairmaire
Compsoglypta perrieri Fairmaire, 1904: 233–234
(orig. desc.); Marie and Lesne, 1917: 30;
Obenberger, 1926: 197; 1928: 339; 1934:
795.
Compsoglypha perrieri: Théry, 1905: 177.
Psiloptera sculpticollis Kerremans, 1911: 347
(orig. desc., nom. superfl. perrieri Fairmaire);
Obenberger, 1926: 197; 1928: 339; 1934:
795.
Genus 6. CORAEBASTUS Fairmaire
Coraebastus Fairmaire, 1896: 456–457 (orig.
desc.); Kerremans, 1903: 223; Théry, 1905:
116; Marie and Lesne, 1917: 31; Obenberger,
1934: 809; Descarpentries, 1978: 107;
Bellamy, 1985: 424.
60
Type species: Coraebastus quinquepustulatus
Fairmaire (original monotypy).
Annietta Obenberger, 1931: 184; 1934: 809;
Descarpentries, 1978: 108 (synonym of
Coraebastus); Bellamy, 1985: 424.
Type species: Annietta imperatrix Obenberger
and the Comoro Islands. While I am in agreement
with Descarpentries (1978) about the generic
synonymy, I do not agree with the species synonymy. I have examined at least one specimen of each
species from Madagascar and the Comores and
find species level distinctions that validate both
taxa. This is not surprising given the different
species in other buprestid genera found in
these two places (e.g. Polybothris Spinola, Chalcophoropsis monochroma from the Comores
(Gianasso 1999)).
12. imperatrix (Obenberger), name reinstated
Annietta imperatrix Obenberger, 1931: 184–185
(orig. desc.); 1934: 809.
Coraebastus imperatrix: Descarpentries, 1978:
108 (syn. quinquepustulatus).
13. quinquepustulatus Fairmaire
Coraebastus quinquepustulatus Fairmaire,
1896: 456 (orig. desc.); Alluaud, 1900: 197;
Kerremans, 1903: 223; Théry, 1905: 116;
Marie and Lesne, 1917: 31; Obenberger,
1934: 809; Descarpentries, 1978: 107.
Genus 7. CORAEBUS Gory and Laporte
Coraebus Gory and Laporte, 1839: 1–2 (orig.
desc.); Gemminger and Harold, 1869: 1429;
Saunders, 1871: 104; Kerremans, 1892: 227;
1893: 117; Théry, 1905: 122; Obenberger,
1934: 817; Bellamy, 1985: 425; Holynski,
1993: 15.
Type species: Buprestis undatus Fabricius (subsequent designation: Descarpentries and
Villiers, 1967: 471).
REMARKS. This genus is widespread in the
Palaearctic and Oriental Regions. Representation in
the Afrotropical fauna was eliminated by the generic
transfers of Bellamy (1988) and the placement of
C. insignis in Coraebus is suspect.
14. insignis Kerremans
Coraebus insignis Kerremans, 1898: 83 (orig.
desc.); Alluaud, 1900: 197; Kerremans, 1903:
233; Théry, 1905: 122; Obenberger, 1934:
830.
Genus 8. DISCODEROPSIS Théry
Discoderopsis Théry, 1930b: 168–169 (orig.
desc.); Obenberger, 1934: 795; Bellamy,
1985: 423.
Type species: Discoderopsis incurvata Théry
and is putatively close to the Afrotropical lineage
which contains Pseudokerremansia and Cupriscobina, both Bellamy and Holm (1985).
15. nigrovirens (Fairmaire)
Discoderes nigrovirens Fairmaire, 1899:
116–117 (orig. desc.); Alluaud, 1900: 197;
Kerremans, 1903: 223; Théry, 1905: 114;
Marie and Lesne, 1917: 40.
Phlocteis nigrovirens: Obenberger, 1928: 341;
1934: 798.
Discoderopsis nigrovirens: Théry, 1936a: 60.
Discoderopsis incurvatus Théry, 1930b:
169–170 (orig. desc.); Obenberger, 1934:
7 9 5 ; T hé r y, 1 9 3 6 a : 6 0 ( sy n o n y m o f
nigrovirens).
Genus 9. ENTOMOGASTER Saunders
Entomogaster Saunders, 1871: 102; Kerremans,
1892: 223; 1893: 122; Fairmaire, 1899:
114–115 (description); Kerremans, 1900:
3 1 9 ; 1 9 0 3 : 2 2 5 ; T hé r y, 1 9 0 5 : 1 1 7 ;
Obenberger, 1928: 340; Théry, 1930b: 168;
Obenberger, 1934: 808; Théry, 1943: 149;
Bellamy, 1985: 424; Holynski, 1993: 15;
Bellamy, 1997: 78.
Type species: Anthaxia sexpunctatus Gory and
Laporte (original monotypy).
Entomoderes Fairmaire, 1903a: 30 (in part,
lapsus calami for Entomogaster); Marie and
Lesne, 1917: 45.
and shows no particular similarities to any lineage of
the Afrotropical fauna. There are many hundreds of
specimens of Entomogaster in the study material in
Paris.
16. aequalipennis Obenberger
Entomogaster aequalipennis Obenberger, 1931:
197–198 (orig. desc.); 1934: 808.
17. amplicollis Fairmaire
Entomogaster amplicollis Fairmaire, 1900:
243–244 (orig. desc.); Kerremans, 1900: 319;
1903: 226; Théry, 1905: 118; Marie and
Lesne, 1917: 46; Obenberger, 1928: 340;
1934: 808.
18. apollo Obenberger
Entomogaster apollo Obenberger, 1946:
160–161 (orig. desc.).
19. asperulus Théry
Entomogaster asperulus Théry, 1905: 120, 179
(orig. desc.), fig. 20; Obenberger, 1928: 340;
1934: 808.
20. carli Théry
Entomogaster carli Théry, 1936b: 697–699 (orig.
desc.), fig. 1.
21. descarpentriesi Théry
Entomogaster descarpentriesi Théry, 1937:
95–96 (orig. desc.), fig. 6.
22. funebris Théry
Entomogaster funebris Théry, 1937: 98–100
23. grandis Théry
Entomogaster grandis Théry, 1937: 100–102
24. griseofasciatus Obenberger
Entomogaster griseofasciatus Obenberger,
1946: 162–163 (orig. desc.).
25. hoschecki Théry
Entomogaster hoschecki Théry, 1937: 96–98
61
(orig. desc.); Obenberger, 1928: 340; 1934:
808.
31. modestus Fairmaire
Entomogaster modestus Fairmaire, 1899:
115–116 (orig. desc.); Alluaud, 1900: 198;
Kerremans, 1900: 319; 1903: 226; Théry,
1905: 118; Marie and Lesne, 1917: 46;
Obenberger, 1928: 340; 1934: 808; Bellamy,
1997: 78.
32. octoguttatus (Kerremans)
Agrilus octoguttatus Kerremans, 1894: 353 (orig.
desc.).
Entomogaster octoguttatus: Fairmaire, 1899:
116; Alluaud, 1900: 198; Kerremans, 1900:
3 1 9 ; 1 9 0 3 : 2 2 5 ; T hé r y, 1 9 0 5 : 1 1 8 ;
Obenberger, 1928: 340; 1934: 809.
33. posticalis (Fairmaire)
Entomoderes posticalis Fairmaire, 1903a: 29
(orig. desc.).
Entomogaster posticalis: Fairmaire, 1903b: 201–
202; Théry, 1905: 119; Marie and Lesne, 1917:
45, 46; Obenberger, 1928: 340; 1934: 809.
34. pumilio Obenberger
Entomogaster pumilio Obenberger, 1946:
163–164 (orig. desc.).
26. iris Obenberger
Entomogaster iris Obenberger, 1946: 160 (orig.
desc.).
35. rufobasalis (Fairmaire)
Cisseis rufobasalis Fairmaire, 1897: 380 (orig.
desc.); Alluaud, 1900: 197.
Entomogaster rufobasalis: Kerremans, 1903:
340; 1934: 809.
27. kerremansi Fairmaire
Entomogaster kerremansi Fairmaire, 1899: 115
(orig. desc.); Alluaud, 1900: 197; Kerremans,
1900: 319; 1903: 225; Théry, 1905: 118; Marie
and Lesne, 1917: 46; Obenberger, 1928: 340;
1934: 808; Bellamy, 1997: 78.
Entomogaster lantzi Théry (nom. nud., teste
Obenberger, 1928: 340); Obenberger, 1934:
808.
36. sexpunctatus (Gory and Laporte)
Anthaxia sexpunctatus Gory and Laporte, 1839:
36–37 (orig. desc.); Gemminger and Harold,
1869: 1391.
Entomogaster sexpunctatus: Saunders, 1871:
102; Kerremans, 1892: 223; 1893: 122;
Alluaud, 1900: 198; Kerremans, 1900: 319;
1928: 340; 1934: 809; Bellamy, 1997: 78.
28. maculipictus Obenberger
Entomogaster maculipictus Obenberger, 1931:
197 (orig. desc.); 1934: 808.
29. mandrakanus Obenberger
Entomogaster mandrakanus Obenberger, 1946:
161–162 (orig. desc.).
Genus 10. EUMORPHOCERUS Théry
Eumorphocerus Théry, 1930b: 166–167 (orig.
d e sc . ) ; O be n be r g e r, 1 9 3 4 : 8 0 9 ;
Descarpentries, 1968: 934–935 (spp. key);
Bellamy, 1985: 424.
Type species: Eumorphocerus laticornis Théry
30. mocquerysi Théry
Entomogaster mocquerysi Théry, 1905: 120
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37. costipennis Descarpentries
Eumorphocerus costipennis Descarpentries,
1968: 935 (orig. desc.).
38. laticornis Théry
Eumorphocerus laticornis Théry, 1930b:
167–168 (orig. desc.); Obenberger, 1934:
809; Descarpentries, 1968: 934.
39. peyrierasi Descarpentries
Eumorphocerus peyrierasi Descarpentries,
1968: 935 (orig. desc.), fig. 15.
40. vadoni Descarpentries
Eumorphocerus vadoni Descarpentries, 1968:
935 (orig. desc.), fig. 14.
Genus 11. EVIMANTIUS Kerremans
Evimantius Kerremans, 1891: clxiii (orig. desc.);
1892: 237; 1893: 118; 1903: 245; Théry, 1905:
123; 1930b: 168; Obenberger, 1934: 810;
Bellamy, 1985: 424; Holynski, 1993: 15;
Bellamy, 1997: 77.
Type species: Evimantius rufopictus Kerremans
(subsequent designation: Bellamy 1997).
Evimantius Deyrolle, 1864: 113 (nom. nud.);
Saunders, 1871: 137.
Type species: Evimantius rufopictus La Férte
and seems close to species of Anaphlocteis
Bellamy in the Afrotropical fauna.
41. curvicollis Kerremans
Evimantius curvicollis Kerremans, 1891: clxiv
(orig. desc.); 1892: 237; Alluaud, 1900: 198;
Kerremans, 1903: 245; Théry, 1905: 124;
Obenberger, 1934: 810; Bellamy, 1997: 77.
42. rufopictus Kerremans
Evimantius rufopictus Kerremans, 1891: clxiv
(orig. desc.); 1892: 237; Alluaud, 1900: 198;
Kerremans, 1903: 245, plate 3, fig. 4; Théry,
1905: 123; Obenberger, 1934: 810; Bellamy,
1997: 77.
Evimantius rufopictus La Férte (nom. nud.) vide
Deyrolle, 1864: 113.
Evimantius olsufiewi Hoscheck, 1931: 733–735
(orig. desc.); Obenberger, 1934: 810;
Bellamy, 1997: 77 (syn. rufopictus).
Evimantius strandi Obenberger, 1931: 181 (orig.
desc.); 1934: 810; Bellamy, 1997: 77 (syn.
rufopictus).
Evimantius embriki Obenberger, 1936: 139 (orig.
desc.); Bellamy, 1997: 77 (syn. rufopictus).
Genus 12. GIGANTOCORAEBUS Obenberger
Gigantocoraebus Obenberger, 1942: 12–13
(orig. desc.); Bellamy, 1985: 425.
Type species: Buprestis lumbaris Klug (original
designation).
REMARKS. This genus is endemic to Madagascar,
quite distinct from other members of the Madagascan and Afrotropical faunas and is not represented
in the Cerceris-collected material.
43. lumbaris (Klug)
Buprestis lumbaris Klug, 1833: 62 (orig. desc.).
Coraebus lumbaris: Gemminger and Harold,
1869: 1431.
Discoderes lumbaris: Saunders, 1871: 109;
Kerremans, 1892: 236; Alluaud, 1900: 197.
Phlocteis lumbaris: Kerremans, 1903: 257;
Théry, 1905: 125; Obenberger, 1934: 798.
Gigantocoraebus lumbaris: Obenberger, 1942:
13, fig. 5.
Genus 13. JAROSLAVIA Obenberger
Jaroslavia Obenberger, 1942: 11–12 (orig.
Type species: Jaroslavia ianthinipennis
REMARKS. This monotypic genus is endemic to
Madagascar.
44. ianthinipennis Obenberger
Jaroslavia ianthinipennis Obenberger, 1942: 12
Genus 14. MANDRITSARIA Obenberger
Mandritsaria Obenberger, 1942: 7–8 (orig.
desc.); Descarpentries, 1968: 932–933 (spp.
key); Bellamy, 1985: 425.
Type species: Amorphosoma inaequale Gory
and Laporte (original designation).
and shows no particular affinity to any group of the
Afrotropical fauna.
45. ambositrensis Descarpentries
Mandritsaria ambositrensis Descarpentries,
1968: 933 (orig. desc.).
46. antamponensis Descarpentries
Mandritsaria antamponensis Descarpentries,
1968: 932 (orig. desc.).
47. catalai Descarpentries
Mandritsaria catalai Descarpentries, 1968: 933
48. inaequalis (Gory and Laporte)
Amorphosoma inaequale Gory and Laporte,
1839: 11–12 (orig. desc.).
Discoderes inaequalis: Gemminger and Harold,
1869: 1433; Saunders, 1871: 109; Kerremans, 1892: 236; 1903: 223; Alluaud, 1900:
196; Théry, 1905: 114.
Phlocteis inaequalis: Obenberger, 1934: 798.
Mandritsaria inaequalis: Obenberger, 1942: 8,
fig. 4; Descarpentries, 1968: 933.
Mandritsaria peyrierasi Descarpentries, 1968:
933 (orig. desc.).
50. sicardi Descarpentries
Mandritsaria sicardi Descarpentries, 1968: 933
(orig. desc.).
Mandritsaria vadoni Descarpentries, 1968: 932
Genus 15. MAROANTSETRA Théry
Maroantsetra Théry, 1937: 102–103 (orig. desc.);
Bellamy, 1985: 425.
Type species: Maroantsetra superba Théry (original monotypy).
63
1905: 118; Marie and Lesne, 1917: 46; Théry,
1930b: 168.
Meliboeus bipustulatus: Obenberger, 1934: 808,
899.
55. cyanipennis (Fairmaire)
Entomogaster cyanipennis Fairmaire, 1903b:
201 (orig. desc.); Théry, 1905: 119; Marie and
Lesne, 1917: 46; Théry, 1930b: 168;
Obenberger, 1928: 340.
Meliboeus cyanipennis: Obenberger, 1934: 808,
901.
56. hovanus Théry
Meliboeus hovanus Théry, 1937: 107 (orig.
desc.), fig. 12.
57. insipidus Théry
Meliboeus insipidus Théry, 1905: 123 (orig.
desc.), fig. 21; Obenberger, 1934: 902.
58. miliaris (Gory and Laporte)
Agrilus miliaris Gory and Laporte, 1839: 32 (orig.
desc.); Gemminger and Harold, 1869: 1442.
Meliboeus miliaris: Saunders, 1871: 105;
Kerremans, 1892: 232; Alluaud, 1900: 198;
Kerremans, 1903: 236; Théry, 1905: 123;
Obenberger, 1934: 903.
Several new species await description in the Paris
museum.
59. vagecostatus Théry
Meliboeus vagecostatus Théry, 1937: 108 (orig.
desc.), fig.13.
52. superba Théry
Maroantsetra superba Théry, 1937: 103–104
Genus 17. MIDONGYA Obenberger
Midongya Obenberger, 1942: 9–10 (orig. desc.);
Bellamy, 1985: 425.
Type species: Midongya mirabilis Obenberger
Genus 16. MELIBOEUS Deyrolle
Meliboeus Deyrolle, 1864: 132 (orig. desc.);
Gemminger and Harold, 1869: 1429;
Saunders, 1871: 107; Obenberger, 1934:
880; Bellamy, 1985: 426.
Type species: Agrilus episcopalis Mannerheim
(subsequent designation: Schaefer 1949:
333).
REMARKS. This very speciose genus is widespread
in the Old World except for Australasia.
53. albomaculatus Théry
Meliboeus albomaculatus Théry, 1937: 108–110
54. bipustulatus (Fairmaire)
Entomogaster bipustulatus Fairmaire, 1901: 161
(orig. desc.); Kerremans, 1903: 226; Théry,
Madagascar and at least one additional species is
present in the Paris collection.
60. mirabilis Obenberger
Midongya mirabilis Obenberger, 1942: 10–11
Genus 18. PARADORA Kerremans
Paradora Kerremans, 1900: 323 (orig. desc.);
1922: 127; 1934: 796 (syn. Phlocteis); 1935a:
87; Bellamy, 1985: 424; 1986: 128; 1991: 5;
1997: 76.
Type species: Buprestis pumicatus Klug (subsequent designation: Bellamy, 1997).
64
and is quite distinct from the Afrotropical fauna
despite any relationship implied by the name
Paradorella Obenberger from South Africa (see
Bellamy, 1989).
Subgenus Paradora (s. str.)
61. impetigiosus (Gory)
Coraebus impetigiosus Gory, 1841: 277–278
(orig. desc.), plate 46, fig. 269.
Discoderes impetigiosus: Kerremans, 1892: 236.
Paradora impetigiosus: Kerremans, 1900: 323;
1903: 248; Obenberger, 1935a: 87; Bellamy,
1997: 76.
62. pumicatus (Klug)
Buprestis pumicatus Klug, 1833: 62 (orig. desc.).
Coroebus pumicatus: Gory and Laporte, 1839:
6; Gemminger and Harold, 1869: 1431.
Discoderes pumicatus: Saunders, 1871: 109;
Kerremans, 1892: 237; Alluaud, 1900: 197.
Paradora pumicatus: Kerremans, 1900: 323;
1934: 799; 1935a: 87; Bellamy, 1997: 76.
63. purpureosignata Hoschek
Paradora purpureosignata Hoschek, 1931:
735–736 (orig. desc.).
Phlocteis purpureosignata: Obenberger, 1934:
799.
64. saxosicollis (Fairmaire)
Discoderes saxosicollis Fairmaire, 1897: 381
(orig. desc.); Alluaud, 1900: 197.
Paradora saxosicollis: Kerremans, 1900: 323;
1903: 249; Théry, 1905: 124; Bellamy, 1997:
76.
Discoderes saxosicollis: Marie and Lesne, 1917:
40.
Phlocteis saxosicollis: Obenberger, 1928: 341;
1934: 799; 1935a: 87.
65. zonata (Kerremans)
Phlocteis zonata Kerremans, 1899: 277–278
(orig. desc.); Alluaud, 1900: 198.
Paradora zonata: Kerremans, 1900: 323; 1903:
799; 1935a: 87; Bellamy, 1997: 76.
115; Marie and Lesne, 1917: 40.
Phlocteis corynthius Obenberger, 1928: 341
(lapsus calami); Bellamy,1997: 77.
Phlocteis corynthia Obenberger, 1934: 796
(lapsus calami); Bellamy, 1997: 77.
Paradora (Speophora) corinthia: Bellamy, 1997:
77.
Paradora (Speophora) seyrigi Théry, 1937:
92–94 (orig. desc.), fig. 5; Bellamy 1997: 77
(syn. corinthia).
Genus 19. PARANASTELLA Obenberger
Paranastella Obenberger, 1931: 198 (orig.
desc.); 1934: 808; Descarpentries, 1984: 57;
Bellamy, 1985: 424
Type species: Paranastella strandi Obenberger
and despite the generic name does not seem
particularly close to Nastella Kerremans from South
Africa (see Bellamy, 1989). A third species has been
located and will be described in the near future.
67. strandi Obenberger
Paranastella strandi Obenberger, 1931: 198–199
(orig. desc.); 1934: 808; Descarpentries,
1984: 58, fig. 3.
68. villiersi Descarpentries
Paranastella villiersi Descarpentries, 1984:
57–58 (orig. desc.), figs 1, 2.
Genus 20. PEYRIERASINA Descarpentries
Peyrierasina Descarpentries, 1975: 22–24 (orig.
Type species: Peyrierasina complanata
Descarpentries (original designation).
Madagascar and quite distinct from its tribal
relatives.
69. complanata Descarpentries
Peyrierasina complanata Descarpentries, 1975:
24–25 (orig. desc.), figs 2, 6–8.
Subgenus Speophora Théry
Paradora (Speophora) Théry, 1937: 91–92 (orig.
desc.); Bellamy, 1985: 424; 1997: 77.
Type species: Paradora (Speophora) seyrigi
Théry (original monotypy).
Genus 21. PHLOCTEIS Kerremans
Phlocteis Kerremans, 1893: 118 (orig. desc.);
1903: 257; Théry, 1905: 10, 125; Obenberger,
1928: 341; 1934: 795; Bellamy, 1985: 423;
1986: 128 (rev., spp. key); 1988: 112; 1991: 5.
Type species: Buprestis exasperata Swartz (original designation).
66. corinthia (Fairmaire)
Discoderes corinthia Fairmaire, 1901: 162 (orig.
REMARKS. This genus is restricted to the Afro-
tropical region and Madagascar (see Bellamy,
1986).
70. hova Théry
Phlocteis hova Théry, 1937: 89–91 (orig. desc.),
fig. 4.
71. humeralis (Waterhouse)
Discoderes humeralis Waterhouse, 1887: 293
(orig. desc.); Kerremans 1892: 236; Alluaud,
1900: 196.
Phlocteis humeralis: Kerremans, 1903: 257;
Théry, 1905: 125; Obenberger, 1928: 341;
1934: 797.
Genus 22. PSEUDAGRILUS Laporte
Pseudagrilus Laporte, 1835: 165 (orig. desc.);
Gory and Laporte, 1839: 1; Deyrolle, 1864:
116; Gemminger and Harold, 1869: 1434;
1893: 119; Théry, 1930a: 122 (rev., spp. key);
Obenberger, 1934: 872; Bellamy, 1985: 426;
1988: 111; 1991: 8; Holynski, 1993: 15.
Type species: Pseudagrilus splendidus Laporte
Entomoderes Fairmaire, 1903a: 30 (in part,
lapsus calami for Entomogaster); Marie and
Lesne, 1917: 45; Obenberger, 1934: 874;
Théry, 1955: 397.
REMARKS. This genus is restricted to the
Afrotropical region and Madagascar.
72. scintillans (Fairmaire)
Entomoderes scintillans Fairmaire, 1903a: 30
(orig. desc.).
Entomogaster scintillans: Fairmaire, 1903b: 201;
Théry, 1905: 119; Marie and Lesne, 1917: 45,
46.
Pseudagrilus scintillans: Obenberger, 1934: 874.
Pseudagrilus scintillans var. madegassus
Obenberger, 1924: 112; 1934: 874 (var.
scintillans). Syn. nov.
Genus 23. PSEUDOCORAEBUS Théry
Pseudocoraebus Théry, 1905: 121 (orig. desc.);
Obenberger, 1934: 813; Bellamy, 1985: 424.
Type species: Pseudocoraebus mocquerysi
Théry (original monotypy).
and is distinct from the Afrotropical fauna.
73. mocquerysi Théry
Pseudocoraebus mocquerysi Théry, 1905:
121–122 (orig. desc.), fig. 18; Obenberger,
1934: 813.
65
74. strandi Obenberger
Pseudocoraebus strandi Obenberger, 1931: 183
(orig. desc.); 1934: 813.
Genus 24. SAMBIRANIA Obenberger
Sambirania Obenberger, 1942: 8 (orig. desc.);
Bellamy, 1985: 425.
Ty p e sp e c i e s: S a m b i r a ni a b r a c h y s o m a
REMARKS. This genus is endemic to Madagascar.
75. aeneicollis Descarpentries
Sambirania aeneicollis Descarpentries, 1968:
930 (orig. desc.).
76. brachysoma Obenberger
Sambirania brachysoma Obenberger, 1942: 8–9
(orig. desc.), figs 1, 6; Descarpentries, 1968:
929, fig. 5.
77. dubia Descarpentries
Sambirania dubia Descarpentries, 1968: 930
78. impedita Descarpentries
Sambirania impedita Descarpentries, 1968: 929
(orig. desc.), figs 4, 8.
79. incerta Descarpentries
Sambirania incerta Descarpentries, 1968: 930
80. inflata Descarpentries
Sambirania inflata Descarpentries, 1968: 929
(orig. desc.), figs 6, 7.
81. plaicollis Descarpentries
Sambirania plaicollis Descarpentries, 1968: 930
(orig. desc.).
82. rotundipennis Descarpentries
Sambirania rotundipennis Descarpentries, 1968:
930 (orig. desc.).
83. suspecta Descarpentries
Sambirania suspecta Descarpentries, 1968: 930
Genus 25. SAMBUS Deyrolle
Sambus Deyrolle, 1864: 210 (orig. desc.);
Gemminger and Harold, 1869: 1434;
1 8 9 3 : 1 1 9 ; O be n be r g e r, 1 9 3 4 : 8 6 4 ;
Bellamy, 1985: 425; 1988: 111; 1991: 8;
66
Holynski, 1993: 15.
Type species: Sambus lafertei Deyrolle (subsequent designation: Toyama, 1986: 58).
REMARKS. This genus is widespread in the Old
World with the exception of the western Palaearctic.
Sambus is allied to Pseudagrilus with the synapomorphy of saltatorial hind legs.
84. douquetteae Descarpentries
Sambus douquetteae Descarpentries, 1968: 928
(orig. desc.).
85. madagascariensis Descarpentries
Sambus madagascariensis Descarpentries,
1968: 928 (orig. desc.).
Sambus peyrierasi Descarpentries, 1968: 927
87. seyrigi Descarpentries
Sambus seyrigi Descarpentries, 1968: 928 (orig.
desc.), fig. 2.
mimicry complex with Chloricala balachowskyi.
91. albopunctata Théry
Seyrigia albopunctata Théry, 1937: 106–107
Genus 28. SUAREZIA Théry
Suarezia Théry, 1912: 119–120 (orig. desc., nom.
praeocc. Suarezius Fairmaire, lapsus
calami); Obenberger, 1922: 127; 1934: 795;
Théry, 1936a: 60; Bellamy, 1985: 423.
Type species: Suarezia albicollis Théry (original
monotypy).
Suarezina Théry, 1936a: 60 (nom. nov. Suarezia
Théry, nom. superfl.); Bellamy, 1985: 423.
and may be distantly related to the Afrotropical
genus Euchroaria Obenberger.
92. albicollis Théry
Suarezia albicollis Théry, 1912: 120–121 (orig.
desc.); Obenberger, 1934: 795.
89a. nodosa Descarpentries
Seranambia nodosa Descarpentries, 1974:
585–586 (orig. desc.), figs 9, 10.
Undefined placement
The following 11 species were originally
described in one of two coraebine genera:
Discoderes Chevrolat or Phlocteis Kerremans.
Discoderes was last discussed by Bellamy (1986)
and is restricted to sub-Saharan Africa, with two
species distributed in habitats directly below the
Sahara. Phlocteis is a small genus with three
sub-Saharan African species also last discussed by
Bellamy (1986). As noted above, two Madagascan
species, P. hova Théry and P. humeralis (Waterhouse), are placed therein. The types of each of
these species require study and comparison to the
currently more restricted definition of these two
genera and the other Madagascan coraebines
before transfers to other genera will be possible.
89b. nodosa ssp. vondrozensis Descarpentries
Seranambia nodosa ssp. vondrozensis
Descarpentries, 1974: 586 (orig. desc.).
93. ambanjana (Obenberger)
Phlocteis ambanjana Obenberger, 1944:
421–422 (orig. desc.).
90. succinicola Descarpentries
Seranambia succinicola Descarpentries, 1974:
586–587 (orig. desc.), fig. 11.
94. ampliata (Fairmaire)
Discoderes ampliata Fairmaire, 1905: 124 (orig.
desc.); Théry, 1905: 178; Marie and Lesne,
1917: 40. Phlocteis ampliata: Obenberger,
1928: 341; 1934: 796.
Sambus vadoni Descarpentries, 1968: 927 (orig.
desc.), fig. 1.
Genus 26. SERANAMBIA Descarpentries
Seranambia Descarpentries, 1974: 584–585
(orig. desc.); Bellamy, 1985: 424.
Type species: Seranambia nodosa Descarpentries (original designation).
REMARKS. This genus is endemic to Madagascar.
Genus 27. SEYRIGIA Théry
Seyrigia Théry, 1937: 104–105 (orig. desc.);
Bellamy, 1985: 424.
Type species: Seyrigia albopunctata Théry (original designation).
and the single species seems to be involved in a
95. cavifrons (Fairmaire)
Discoderes cavifrons Fairmaire, 1904: 234–235
(orig. desc.); Théry, 1905: 178; Marie and
Lesne, 1917: 40.
Phlocteis cavifrons: Obenberger, 1928: 341;
1934: 796.
96. goudotii (Thomson)
Discoderes goudotii Thomson, 1878: 82–83
(orig. desc.); Kerremans, 1885: 151; 1892:
236; 1903: 223; Alluaud, 1900: 196; Théry,
1905: 114; 1936a: 60.
Discoderes goudoti (lapsus calami): Kerremans,
1885: 151; 1892: 236; 1903: 223; Alluaud,
1900: 196; Théry, 1905: 114; 1936a: 60.
Phlocteis goudoti: Obenberger, 1934: 797.
97. iridipennis (Obenberger)
Phlocteis iridipennis Obenberger, 1944:
420–421 (orig. desc.).
98. mucorea (Fairmaire)
Discoderes mucorea Fairmaire, 1902a: 80 (orig.
Phlocteis mucorea: Obenberger, 1928: 341;
1934: 798.
REMARKS. Obenberger (1928) identified this
species as one warranting a ‘new genus’ in a brief
discussion about the genera Discoderes and
Phlocteis.
99. ochraceopicta (Fairmaire)
Discoderes ochraceopicta Fairmaire, 1888: 56
(orig. desc.); Kerremans, 1892: 237; Alluaud,
1900: 197; Kerremans, 1903: 223; Théry,
1905: 114, fig. 19; Marie and Lesne, 1917: 40;
Théry, 1937: 98.
Phlocteis ochraceopicta: Obenberger, 1934: 798.
100. perrieri (Fairmaire)
Discoderes perrieri Fairmaire, 1900: 244 (orig.
Phlocteis perrieri: Obenberger, 1928: 341; 1934:
798.
101. polychroa (Fairmaire)
Discoderes polychroa Fairmaire, 1904: 234
(orig. desc.); Théry, 1905: 177; Marie and
Lesne, 1917: 40.
Phlocteis polychroa: Obenberger, 1928: 341;
1934: 798.
102. scriptus (Théry)
Discoderes scriptus Théry, 1937: 85–86 (orig.
desc.), fig. 2.
103. simplicifrons (Fairmaire)
Discoderes simplicifrons Fairmaire, 1901: 163
(orig. desc.); Kerremans, 1903: 223; Théry,
1905: 115; Marie and Lesne, 1917: 40.
Phlocteis simplicifrons: Obenberger, 1934: 799.
67
Doubtful taxa
Anadora hova Kerremans, 1899: 277 (orig.
desc.); Alluaud, 1900: 197; Kerremans, 1903:
2 5 8 ; T hé r y, 1 9 0 5 : 1 2 5 ; 1 9 2 7 : 2 6 0 ;
Obenberger, 1934: 794 (howa, lapsus
calami).
REMARKS. Anadora Kerremans is another endemic
Subsaharan African genus last revised by Bellamy
(1986). Théry (1905) listed this taxon as a member
of the Madagascan fauna and later (Théry 1927)
synonymized it with A. mechowi Quedenfeldt.
Obenberger (1934), however, listed it as a valid
species in his catalogue. Since there are no other
specimens of Anadora in the vast Madagascan
coraebine collection in the Paris museum, I support
the synonymy and do not consider either the genus
or species as valid members of the Madagascan
fauna.
In the Paris museum general collection and in the
Obenberger collection in Prague, there are a number of Madagascan coraebines labelled as ‘types’
of various genera and species. These are taxa
a p p a r e n t l y i n t e n d e d f o r d e sc r i p t i o n by
Descarpentries, Théry (both Paris collection), or
Obenberger, and thus are simply manuscript or ‘in
collection’ names. None of these names appear to
be valid as I cannot find descriptions or any mention
in the literature or the Zoological Record. I will not list
any of these names here as that would add confusion to the discussion and nomina nuda to the literature. Should any of these names be found in the
literature, they will be discussed in a forthcoming
contribution to this series.
ACKNOWLEDGEMENTS
For access to the collections under their respective care or for the loan of specimens, I thank J. J.
Menier, Museum National d’Histoire Naturelle,
Paris, France; S. Bílý, Narodni Museum, Prague,
Czech Republic; J. Cools, Institut Royal des
Sciences Naturelles de Belgique, Brussels,
Belgium; R. Holynski, Milanowek, Poland; M. Kerley,
The Natural History Museum, London, United Kingdom; T. Neef de Sainval, Brussels; E. Sprecker,
Naturhistorisches Museum, Basel, Switzerland and
M. Uhlig, Museum für Naturkunde der Humboldt
Universität, Berlin, Germany. I also thank the late A.
Descarpentries for inviting me to join him in working
on this great treasure of beautiful beetles back in
1988; sorry it took so long to get started! Lastly to
Ted MacRae, Monsanto Corporation, St Louis,
Missouri and Rick Westcott, Oregon Department of
Agriculture, Salem, Oregon, my appreciation for
their reading of the original draft and for suggestions toward its improvement.
68
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NOTE ADDED IN PROOF
A new paper by Kuban et al. (2001) was received after this paper was in proof stage. The authors recognized Coraebini at the tribal rank and proposed a phylogenetic arrangement of ten subtribes. The entire
compliment of Madagascan genera were placed in the new subtribe Toxoscelina Majer.
Reference
Kubán,
$ V., Majer, K. and Kolibá, J., 2001. Classification of the tribe Coraebini Bedel, 1921 (Coleoptera, Buprestidae,
Agrilinae). Acta Musei Moraviae, Scientiae biologicae (Brno) 85 (2000): 185–287.

The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae

Transcription

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