The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae
Transcription
The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae
The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae: Agrilinae: Agrilini): introduction and systematic catalogue* C. L. Bellamy† Transvaal Museum, P.O. Box 413, Pretoria, 0001 South Africa Bellamy, C. L., 2001. The Madagascan Coraebina Bedel, 1921 (Coleoptera: Buprestidae: Agrilinae: Agrilini): introduction and systematic catalogue. Annals of the Transvaal Museum 38: 57–70. An introduction to a vast descriptive and systematic project that will consider the large fauna of coraebine jewel beetles from Madagascar and the Comoro Islands is presented. The described species are summarized in a catalogue that contains 28 genera, one subgenus, 103 species and one subspecies. Coraebastus imperatrix (Obenberger), from Madagascar, is resurrected from synonymy under C. quinquepustulatus Fairmaire, from the Comoro Islands. Chloricala gratiosa Kerremans var. octosticta Obenberger and var. impunctata Théry and Pseudagrilus scintillans var. madegassus Obenberger are synonymized under their nominate species. Le vaste projet de description systématique de l’importante faune des Buprestes Coraebinae de Madagascar et des Comores est présenté. La description des espèces est résumée dans un catalogue contenant 28 genres, un sous-genre, 103 espèces et une sous-espèce. Coraebastus imperatrix (Obenberger), de Madagascar, est rétabli au rang d’espèce. Il n’est plus synonyme de C. quinquepustulatus Fairmaire des Comores. Les var. otosticta Obenberger et impunctata Théry du Chloricala gratiosa Kerremans ainsi que le Pseudagrillus scintillans var. madegassus Obenberger sont synonymisés sous le nom de leurs espèces nominales. Keywords/Mots Clés: Coleoptera, Buprestidae, Agrilinae, Coraebina, Madagascar. INTRODUCTION There are many accounts of a proliferation of species in the literature. Perhaps these come more frequently now with more attention being paid to inventorying threatened habitats as world populations rise and development invades once pristine tracts of nature. In my experience, there is nothing that compares to the vast and largely undescribed jewel beetle subtribe Coraebina Bedel (1921) (Coleoptera: Buprestidae: Agrilinae: Agrilini) from Madagascar. This is the second contribution in a series that is anticipated to take many years to complete; the first was by Bellamy (1997). During the latter half of the 20th century, French and Malagasy entomologists discovered that nest-provisioning wasps (Hymenoptera: Sphecoidea: Philanthidae: Cerceris Latreille) were using small jewel beetles as their primary prey. The two principal collectors on the majority of specimen data labels are P. Vadon and A. Peyrieras. The beetles are paralyzed by the sting of female wasps and then carried back to the nests, whereupon an egg is laid and the emerging larva feeds on the beetle before pupating inside the hollow body. By following the wasps and learning to recognize their burrows *Third contribution to the knowledge of Madagascar Buprestidae. † Present address: Entomology Section, Natural History Museum of Los Angeles County, 900 Exposition Blvd., Los Angeles, California, 90007, U.S.A. E-mail: [email protected] or nests, these entomologists were able to dig up many comatose beetles, which lay in perfect or near-perfect condition, and return them for preparation and permanent museum storage. The predator–prey relationship between certain species of Cerceris and buprestids has been widely reported for various parts of the world (e.g. in general: Cartwright, 1931; Inglebert, 1993; Scullen and Wold, 1969; Trinidad: Fisher, 1944; Callan, 1990; North America: Grossbeck, 1912; Linsley and MacSwain, 1956; Evans and Rubinik, 1978; Kurczewski and Miller, 1984; Hook and Evans, 1991; Japan: Ohmomo and Usuba, 1997). Several colleagues (W. F. Barr, H. A. Hespenheide, S. G. Wellso, pers. comm.) had told me about the collection of Madagascan buprestids seen in the Museum National d’Histoire Naturelle, Paris, during their respective research visits in the 1970s and 1980s, but until I experienced this firsthand during my initial visit to the museum in late 1995, I really could not understand the breadth of this collection. There are thousands of specimens, prepared and labelled, representing a large number of mostly undescribed species. From an initial attempt to sort and arrange these specimens by the former head of the Coleoptera Department, the late A. Descarpentries, an estimate of the size of this fauna is between 500–600 undescribed species, which may require 20–30 new genera to contain 58 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 them. It seems clear from the literature, as well as from the examples of other buprestid prey in this material, that the Cerceris predators are not particular to coraebine buprestids, nor possibly even just buprestids. There are large series of other buprestid species (e.g., Sponsor Gory and Laporte, see Descarpentries, 1965; Anthaxia Eschscholtz, see Descarpentries, 1967) that are each of about the same general size and shape as the many coraebine species and this suggests that the predators are perhaps more selective about the physical characteristics of the prey, perhaps to be able to contain them within specific morphological adaptations to maintain a hold. Krombein (1981: 8) correlated structure and function when he described and figured the ‘buprestid clamp’ found in members of the ‘bupresticida group’ of Cerceris, including species from Sri Lanka, Europe and North America. The present paper is intended as an introduction to the many nomenclatural acts and descriptive papers that must follow. By introducing the subject and providing an accounting of the current systematics of the known Madagascan coraebine fauna, all subsequent efforts can focus on the description of new taxa and several of the large genera which have never been revised, e.g. Entomogaster Saunders. The catalogue included herein is presented in full, indicating all taxa and the nomenclatural history for each species. This effort to present a full catalogue of the previously described taxa of this subtribe is undertaken now to bring together all of the available information on a large group of species that have mostly appeared in random and disparate descriptive publications and earlier more general catalogues. The chronology of catalogues that have treated these taxa, in part, are: Gemminger and Harold, 1869; Saunders, 1871; Kerremans, 1885, 1892; Alluaud, 1900; Kerremans, 1903; Théry, 1905; Marie and Lesne, 1917; Obenberger, 1934; and Bellamy, 1985. The work of Obenberger (1934) was the fourth of six fascicles of the two volumes on Buprestidae in the Coleopterorum Catalogue, the last effort to catalogue the world’s beetle fauna. Many of the taxa of Madagascan Coraebina were described subsequent to that catalogue and have not been found in the literature after their respective original descriptions. For future geographical notes, distributional accounts and attempts to summarize the range of particular taxa within the Madagascan habitats, I note here that a great many of the older taxa of this group were described from a type locality no more precise than ‘Madagascar’. However, considering the wealth of specimens of many of these taxa that were harvested, some detail will be filled in as species are described and genera are revised. As a reference for Madagascan place names, the work of Viette (1991) will be the standard. The described complement of these beetles is 28 genera, one subgenus, 103 species and one subspecies. The single related taxon, Coraebastus quinquepustulatus Fairmaire, from the Comoro Islands is included with the reinstatement from synonymy of Coraebastus imperatrix (Obenberger) from Madagascar. In the catalogue, the genera and species are listed alphabetically. With so much descriptive work to be done, phylogenetic predictions are premature, since each subsequent descriptive paper will alter the conclusions about transformation character state series and synapomorphies. At the end of the catalogue are listed 11 species names that were originally described in one of two older coraebine genera. These are listed as ‘undefined placement’ pending better understanding of relationships with the many new species amongst which some of these will undoubtedly fit. Following current trends in dealing with infrasubspecific names, I will propose all varietal names listed below as new synonyms (see Bellamy & Peterson, 2000). Abbreviations used herein are: nom. nov. = nomen novum; nom. nud. = nomen nudum; nom. superfl. = superfluous name, i.e. a replacement name that was unnecessary; orig. desc. = original description; rev. = revision; spp. key = species key; syn. = synonym; syn. nov. = new synonym; unvn. = unavailable name, i.e. infrasubspecific names, nomina nuda; var. = variety, variation. SYSTEMATIC CATALOGUE OF THE COROEBINA OF THE MADAGASCAN REGION Genus 1. ANGATRA Descarpentries Angatra Descarpentries, 1969: 599–600 (orig. desc.); Bellamy, 1985: 424. Type species: Angatra magna Descarpentries (original designation). REMARKS. This genus is endemic to Madagascar and apparently has no close relationships to the Afrotropical fauna. 1. magna Descarpentries Angatra magna Descarpentries, 1969: 600–601 (orig. desc.), fig. 1. 2. parva Descarpentries Angatra parva Descarpentries, 1969: 601–602 (orig. desc.), fig. 2. Genus 2. ANTANAMBIA Descarpentries Antanambia Descarpentries, 1975: 19–20 (orig. MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE desc.); Bellamy, 1985: 425. Ty p e s p e c ie s : A n t a n a m b i a n i g r i c a ns Descarpentries (original designation). REMARKS. This genus is endemic to Madagascar and it is not particularly close to any coraebine lineage in sub-Saharan Africa. 3. nigricans Descarpentries Antanambia nigricans Descarpentries, 1975: 21–22 (orig. desc.), figs 1, 3–5. Genus 3. BRACHYDORA Obenberger Brachydora Obenberger, 1923: 30–31 (orig. desc.); 1934: 794; Bellamy, 1985: 423. Ty p e s p e c ie s : B r a c h y d or a m o ns t r um Obenberger (original designation). REMARKS. This genus is endemic to Madagascar and is quite distinct from the Afrotropical fauna. 4. crassa Théry Brachydora crassa Théry, 1937: 87–89 (orig. desc.), fig. 3. 5. deformis (Fairmaire) Discoderes deformis Fairmaire, 1901: 163 (orig. desc.); Kerremans, 1903: 223; Marie and Lesne, 1917: 40. Paradora deformis: Théry, 1905: 125. Brachydora deformis: Obenberger, 1923: 32; 1934: 795. 6. granulum (Fairmaire) Discoderes granulum Fairmaire, 1902b: 350–351 (orig. desc.); Théry, 1905: 116; Marie and Lesne, 1917: 40 Brachydora granulum: Obenberger, 1928: 342; 1934: 795. 7. monstrum Obenberger Brachydora monstrum Obenberger, 1923: 31–32 (orig. desc.); 1934: 795. 8. sicardi (Théry) Paradora sicardi Théry, 1912: 118–119 (orig. desc.,) Brachydora sicardi: Obenberger, 1923: 31; 1934: 795. Genus 4. CHLORICALA Kerremans Chloricala Kerremans, 1893: 119–120 (orig. desc.); Théry, 1905: 120; Bellamy, 1985: 424. Type species: Chloricala gratiosa Kerremans (original monotypy). Chloricalla (lapsus calami): Kerremans, 1900: 319; 1903: 231. 59 REMARKS. This genus is endemic to Madagascar and shows no close similarities to the Afrotropical fauna. The two varietal names deserve no recognition and are technically invalid (ICZN 1999, Article 45.5, 45.6). 9. balachowskyi Descarpentries Chloricala balachowskyi Descarpentries, 1974: 582–584 (orig. desc.), figs 7, 8. 10. gratiosa Kerremans Chloricala gratiosa Kerremans, 1893: 120 (orig. desc.); 1900: 319; 1903: 231, plate 2, fig. 11; Théry, 1905: 120; Obenberger, 1934: 813; 1935b: 20; Théry, 1936a: 59. Chloricala gratiosa var. octosticta Obenberger, 1935b: 20 (orig. desc.); 1934: 813; Théry, 1936a: 59. Syn. nov. Chloricala gratiosa var. impunctata Théry, 1936a: 60–61 (orig. desc.). Syn. nov. Genus 5. COMPSOGLYPTA Fairmaire Compsoglypta Fairmaire, 1904: 233 (orig. desc.); Kerremans, 1910: 3 (syn. Psiloptera), 347; Marie and Lesne, 1917: 30; Obenberger, 1928: 339; 1934: 795; Bellamy, 1985: 423. Type species: Compsoglypta perrieri Fairmaire (original monotypy). Compsoglypha (lapsus calami): Théry, 1905: 177; Obenberger, 1934: 75; Bellamy, 1985: 423. REMARKS. This genus is endemic to Madagascar and its placement in Psilopterina or Coraebina has been somewhat confused as indicated by the synonymy above. I have examined the types of the Fairmaire taxon and am confident that this is a member of the coraebine lineage, albeit very divergent. 11. perrieri Fairmaire Compsoglypta perrieri Fairmaire, 1904: 233–234 (orig. desc.); Marie and Lesne, 1917: 30; Obenberger, 1926: 197; 1928: 339; 1934: 795. Compsoglypha perrieri: Théry, 1905: 177. Psiloptera sculpticollis Kerremans, 1911: 347 (orig. desc., nom. superfl. perrieri Fairmaire); Obenberger, 1926: 197; 1928: 339; 1934: 795. Genus 6. CORAEBASTUS Fairmaire Coraebastus Fairmaire, 1896: 456–457 (orig. desc.); Kerremans, 1903: 223; Théry, 1905: 116; Marie and Lesne, 1917: 31; Obenberger, 1934: 809; Descarpentries, 1978: 107; Bellamy, 1985: 424. 60 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 Type species: Coraebastus quinquepustulatus Fairmaire (original monotypy). Annietta Obenberger, 1931: 184; 1934: 809; Descarpentries, 1978: 108 (synonym of Coraebastus); Bellamy, 1985: 424. Type species: Annietta imperatrix Obenberger (original designation). REMARKS. This genus is endemic to Madagascar and the Comoro Islands. While I am in agreement with Descarpentries (1978) about the generic synonymy, I do not agree with the species synonymy. I have examined at least one specimen of each species from Madagascar and the Comores and find species level distinctions that validate both taxa. This is not surprising given the different species in other buprestid genera found in these two places (e.g. Polybothris Spinola, Chalcophoropsis monochroma from the Comores (Gianasso 1999)). 12. imperatrix (Obenberger), name reinstated Annietta imperatrix Obenberger, 1931: 184–185 (orig. desc.); 1934: 809. Coraebastus imperatrix: Descarpentries, 1978: 108 (syn. quinquepustulatus). 13. quinquepustulatus Fairmaire Coraebastus quinquepustulatus Fairmaire, 1896: 456 (orig. desc.); Alluaud, 1900: 197; Kerremans, 1903: 223; Théry, 1905: 116; Marie and Lesne, 1917: 31; Obenberger, 1934: 809; Descarpentries, 1978: 107. Genus 7. CORAEBUS Gory and Laporte Coraebus Gory and Laporte, 1839: 1–2 (orig. desc.); Gemminger and Harold, 1869: 1429; Saunders, 1871: 104; Kerremans, 1892: 227; 1893: 117; Théry, 1905: 122; Obenberger, 1934: 817; Bellamy, 1985: 425; Holynski, 1993: 15. Type species: Buprestis undatus Fabricius (subsequent designation: Descarpentries and Villiers, 1967: 471). REMARKS. This genus is widespread in the Palaearctic and Oriental Regions. Representation in the Afrotropical fauna was eliminated by the generic transfers of Bellamy (1988) and the placement of C. insignis in Coraebus is suspect. 14. insignis Kerremans Coraebus insignis Kerremans, 1898: 83 (orig. desc.); Alluaud, 1900: 197; Kerremans, 1903: 233; Théry, 1905: 122; Obenberger, 1934: 830. Genus 8. DISCODEROPSIS Théry Discoderopsis Théry, 1930b: 168–169 (orig. desc.); Obenberger, 1934: 795; Bellamy, 1985: 423. Type species: Discoderopsis incurvata Théry (original monotypy). REMARKS. This genus is endemic to Madagascar and is putatively close to the Afrotropical lineage which contains Pseudokerremansia and Cupriscobina, both Bellamy and Holm (1985). 15. nigrovirens (Fairmaire) Discoderes nigrovirens Fairmaire, 1899: 116–117 (orig. desc.); Alluaud, 1900: 197; Kerremans, 1903: 223; Théry, 1905: 114; Marie and Lesne, 1917: 40. Phlocteis nigrovirens: Obenberger, 1928: 341; 1934: 798. Discoderopsis nigrovirens: Théry, 1936a: 60. Discoderopsis incurvatus Théry, 1930b: 169–170 (orig. desc.); Obenberger, 1934: 7 9 5 ; T hé r y, 1 9 3 6 a : 6 0 ( sy n o n y m o f nigrovirens). Genus 9. ENTOMOGASTER Saunders Entomogaster Saunders, 1871: 102; Kerremans, 1892: 223; 1893: 122; Fairmaire, 1899: 114–115 (description); Kerremans, 1900: 3 1 9 ; 1 9 0 3 : 2 2 5 ; T hé r y, 1 9 0 5 : 1 1 7 ; Obenberger, 1928: 340; Théry, 1930b: 168; Obenberger, 1934: 808; Théry, 1943: 149; Bellamy, 1985: 424; Holynski, 1993: 15; Bellamy, 1997: 78. Type species: Anthaxia sexpunctatus Gory and Laporte (original monotypy). Entomoderes Fairmaire, 1903a: 30 (in part, lapsus calami for Entomogaster); Marie and Lesne, 1917: 45. REMARKS. This genus is endemic to Madagascar and shows no particular similarities to any lineage of the Afrotropical fauna. There are many hundreds of specimens of Entomogaster in the study material in Paris. 16. aequalipennis Obenberger Entomogaster aequalipennis Obenberger, 1931: 197–198 (orig. desc.); 1934: 808. 17. amplicollis Fairmaire Entomogaster amplicollis Fairmaire, 1900: 243–244 (orig. desc.); Kerremans, 1900: 319; 1903: 226; Théry, 1905: 118; Marie and Lesne, 1917: 46; Obenberger, 1928: 340; 1934: 808. MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE 18. apollo Obenberger Entomogaster apollo Obenberger, 1946: 160–161 (orig. desc.). 19. asperulus Théry Entomogaster asperulus Théry, 1905: 120, 179 (orig. desc.), fig. 20; Obenberger, 1928: 340; 1934: 808. 20. carli Théry Entomogaster carli Théry, 1936b: 697–699 (orig. desc.), fig. 1. 21. descarpentriesi Théry Entomogaster descarpentriesi Théry, 1937: 95–96 (orig. desc.), fig. 6. 22. funebris Théry Entomogaster funebris Théry, 1937: 98–100 (orig. desc.), fig. 8. 23. grandis Théry Entomogaster grandis Théry, 1937: 100–102 (orig. desc.), fig. 9. 24. griseofasciatus Obenberger Entomogaster griseofasciatus Obenberger, 1946: 162–163 (orig. desc.). 25. hoschecki Théry Entomogaster hoschecki Théry, 1937: 96–98 (orig. desc.), fig. 7. 61 (orig. desc.); Obenberger, 1928: 340; 1934: 808. 31. modestus Fairmaire Entomogaster modestus Fairmaire, 1899: 115–116 (orig. desc.); Alluaud, 1900: 198; Kerremans, 1900: 319; 1903: 226; Théry, 1905: 118; Marie and Lesne, 1917: 46; Obenberger, 1928: 340; 1934: 808; Bellamy, 1997: 78. 32. octoguttatus (Kerremans) Agrilus octoguttatus Kerremans, 1894: 353 (orig. desc.). Entomogaster octoguttatus: Fairmaire, 1899: 116; Alluaud, 1900: 198; Kerremans, 1900: 3 1 9 ; 1 9 0 3 : 2 2 5 ; T hé r y, 1 9 0 5 : 1 1 8 ; Obenberger, 1928: 340; 1934: 809. 33. posticalis (Fairmaire) Entomoderes posticalis Fairmaire, 1903a: 29 (orig. desc.). Entomogaster posticalis: Fairmaire, 1903b: 201– 202; Théry, 1905: 119; Marie and Lesne, 1917: 45, 46; Obenberger, 1928: 340; 1934: 809. 34. pumilio Obenberger Entomogaster pumilio Obenberger, 1946: 163–164 (orig. desc.). 26. iris Obenberger Entomogaster iris Obenberger, 1946: 160 (orig. desc.). 35. rufobasalis (Fairmaire) Cisseis rufobasalis Fairmaire, 1897: 380 (orig. desc.); Alluaud, 1900: 197. Entomogaster rufobasalis: Kerremans, 1903: 226; Théry, 1905: 118; Obenberger, 1928: 340; 1934: 809. 27. kerremansi Fairmaire Entomogaster kerremansi Fairmaire, 1899: 115 (orig. desc.); Alluaud, 1900: 197; Kerremans, 1900: 319; 1903: 225; Théry, 1905: 118; Marie and Lesne, 1917: 46; Obenberger, 1928: 340; 1934: 808; Bellamy, 1997: 78. Entomogaster lantzi Théry (nom. nud., teste Obenberger, 1928: 340); Obenberger, 1934: 808. 36. sexpunctatus (Gory and Laporte) Anthaxia sexpunctatus Gory and Laporte, 1839: 36–37 (orig. desc.); Gemminger and Harold, 1869: 1391. Entomogaster sexpunctatus: Saunders, 1871: 102; Kerremans, 1892: 223; 1893: 122; Alluaud, 1900: 198; Kerremans, 1900: 319; 1903: 226; Théry, 1905: 119; Obenberger, 1928: 340; 1934: 809; Bellamy, 1997: 78. 28. maculipictus Obenberger Entomogaster maculipictus Obenberger, 1931: 197 (orig. desc.); 1934: 808. 29. mandrakanus Obenberger Entomogaster mandrakanus Obenberger, 1946: 161–162 (orig. desc.). Genus 10. EUMORPHOCERUS Théry Eumorphocerus Théry, 1930b: 166–167 (orig. d e sc . ) ; O be n be r g e r, 1 9 3 4 : 8 0 9 ; Descarpentries, 1968: 934–935 (spp. key); Bellamy, 1985: 424. Type species: Eumorphocerus laticornis Théry (original monotypy). 30. mocquerysi Théry Entomogaster mocquerysi Théry, 1905: 120 REMARKS. This genus is endemic to Madagascar and is quite distinct from the Afrotropical fauna. 62 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 37. costipennis Descarpentries Eumorphocerus costipennis Descarpentries, 1968: 935 (orig. desc.). 38. laticornis Théry Eumorphocerus laticornis Théry, 1930b: 167–168 (orig. desc.); Obenberger, 1934: 809; Descarpentries, 1968: 934. 39. peyrierasi Descarpentries Eumorphocerus peyrierasi Descarpentries, 1968: 935 (orig. desc.), fig. 15. 40. vadoni Descarpentries Eumorphocerus vadoni Descarpentries, 1968: 935 (orig. desc.), fig. 14. Genus 11. EVIMANTIUS Kerremans Evimantius Kerremans, 1891: clxiii (orig. desc.); 1892: 237; 1893: 118; 1903: 245; Théry, 1905: 123; 1930b: 168; Obenberger, 1934: 810; Bellamy, 1985: 424; Holynski, 1993: 15; Bellamy, 1997: 77. Type species: Evimantius rufopictus Kerremans (subsequent designation: Bellamy 1997). Evimantius Deyrolle, 1864: 113 (nom. nud.); Saunders, 1871: 137. Type species: Evimantius rufopictus La Férte (original designation). REMARKS. This genus is endemic to Madagascar and seems close to species of Anaphlocteis Bellamy in the Afrotropical fauna. 41. curvicollis Kerremans Evimantius curvicollis Kerremans, 1891: clxiv (orig. desc.); 1892: 237; Alluaud, 1900: 198; Kerremans, 1903: 245; Théry, 1905: 124; Obenberger, 1934: 810; Bellamy, 1997: 77. 42. rufopictus Kerremans Evimantius rufopictus Kerremans, 1891: clxiv (orig. desc.); 1892: 237; Alluaud, 1900: 198; Kerremans, 1903: 245, plate 3, fig. 4; Théry, 1905: 123; Obenberger, 1934: 810; Bellamy, 1997: 77. Evimantius rufopictus La Férte (nom. nud.) vide Deyrolle, 1864: 113. Evimantius olsufiewi Hoscheck, 1931: 733–735 (orig. desc.); Obenberger, 1934: 810; Bellamy, 1997: 77 (syn. rufopictus). Evimantius strandi Obenberger, 1931: 181 (orig. desc.); 1934: 810; Bellamy, 1997: 77 (syn. rufopictus). Evimantius embriki Obenberger, 1936: 139 (orig. desc.); Bellamy, 1997: 77 (syn. rufopictus). Genus 12. GIGANTOCORAEBUS Obenberger Gigantocoraebus Obenberger, 1942: 12–13 (orig. desc.); Bellamy, 1985: 425. Type species: Buprestis lumbaris Klug (original designation). REMARKS. This genus is endemic to Madagascar, quite distinct from other members of the Madagascan and Afrotropical faunas and is not represented in the Cerceris-collected material. 43. lumbaris (Klug) Buprestis lumbaris Klug, 1833: 62 (orig. desc.). Coraebus lumbaris: Gemminger and Harold, 1869: 1431. Discoderes lumbaris: Saunders, 1871: 109; Kerremans, 1892: 236; Alluaud, 1900: 197. Phlocteis lumbaris: Kerremans, 1903: 257; Théry, 1905: 125; Obenberger, 1934: 798. Gigantocoraebus lumbaris: Obenberger, 1942: 13, fig. 5. Genus 13. JAROSLAVIA Obenberger Jaroslavia Obenberger, 1942: 11–12 (orig. desc.); Bellamy, 1985: 424. Type species: Jaroslavia ianthinipennis Obenberger (original designation). REMARKS. This monotypic genus is endemic to Madagascar. 44. ianthinipennis Obenberger Jaroslavia ianthinipennis Obenberger, 1942: 12 (orig. desc.), fig. 3. Genus 14. MANDRITSARIA Obenberger Mandritsaria Obenberger, 1942: 7–8 (orig. desc.); Descarpentries, 1968: 932–933 (spp. key); Bellamy, 1985: 425. Type species: Amorphosoma inaequale Gory and Laporte (original designation). REMARKS. This genus is endemic to Madagascar and shows no particular affinity to any group of the Afrotropical fauna. 45. ambositrensis Descarpentries Mandritsaria ambositrensis Descarpentries, 1968: 933 (orig. desc.). 46. antamponensis Descarpentries Mandritsaria antamponensis Descarpentries, 1968: 932 (orig. desc.). 47. catalai Descarpentries Mandritsaria catalai Descarpentries, 1968: 933 (orig. desc.), fig. 13. MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE 48. inaequalis (Gory and Laporte) Amorphosoma inaequale Gory and Laporte, 1839: 11–12 (orig. desc.). Discoderes inaequalis: Gemminger and Harold, 1869: 1433; Saunders, 1871: 109; Kerremans, 1892: 236; 1903: 223; Alluaud, 1900: 196; Théry, 1905: 114. Phlocteis inaequalis: Obenberger, 1934: 798. Mandritsaria inaequalis: Obenberger, 1942: 8, fig. 4; Descarpentries, 1968: 933. 49. peyrierasi Descarpentries Mandritsaria peyrierasi Descarpentries, 1968: 933 (orig. desc.). 50. sicardi Descarpentries Mandritsaria sicardi Descarpentries, 1968: 933 (orig. desc.). 51. vadoni Descarpentries Mandritsaria vadoni Descarpentries, 1968: 932 (orig. desc.), fig. 12. Genus 15. MAROANTSETRA Théry Maroantsetra Théry, 1937: 102–103 (orig. desc.); Bellamy, 1985: 425. Type species: Maroantsetra superba Théry (original monotypy). 63 1905: 118; Marie and Lesne, 1917: 46; Théry, 1930b: 168. Meliboeus bipustulatus: Obenberger, 1934: 808, 899. 55. cyanipennis (Fairmaire) Entomogaster cyanipennis Fairmaire, 1903b: 201 (orig. desc.); Théry, 1905: 119; Marie and Lesne, 1917: 46; Théry, 1930b: 168; Obenberger, 1928: 340. Meliboeus cyanipennis: Obenberger, 1934: 808, 901. 56. hovanus Théry Meliboeus hovanus Théry, 1937: 107 (orig. desc.), fig. 12. 57. insipidus Théry Meliboeus insipidus Théry, 1905: 123 (orig. desc.), fig. 21; Obenberger, 1934: 902. 58. miliaris (Gory and Laporte) Agrilus miliaris Gory and Laporte, 1839: 32 (orig. desc.); Gemminger and Harold, 1869: 1442. Meliboeus miliaris: Saunders, 1871: 105; Kerremans, 1892: 232; Alluaud, 1900: 198; Kerremans, 1903: 236; Théry, 1905: 123; Obenberger, 1934: 903. REMARKS. This genus is endemic to Madagascar and is quite distinct from the Afrotropical fauna. Several new species await description in the Paris museum. 59. vagecostatus Théry Meliboeus vagecostatus Théry, 1937: 108 (orig. desc.), fig.13. 52. superba Théry Maroantsetra superba Théry, 1937: 103–104 (orig. desc.), fig. 10. Genus 17. MIDONGYA Obenberger Midongya Obenberger, 1942: 9–10 (orig. desc.); Bellamy, 1985: 425. Type species: Midongya mirabilis Obenberger (original designation). Genus 16. MELIBOEUS Deyrolle Meliboeus Deyrolle, 1864: 132 (orig. desc.); Gemminger and Harold, 1869: 1429; Saunders, 1871: 107; Obenberger, 1934: 880; Bellamy, 1985: 426. Type species: Agrilus episcopalis Mannerheim (subsequent designation: Schaefer 1949: 333). REMARKS. This very speciose genus is widespread in the Old World except for Australasia. 53. albomaculatus Théry Meliboeus albomaculatus Théry, 1937: 108–110 (orig. desc.), fig. 14. 54. bipustulatus (Fairmaire) Entomogaster bipustulatus Fairmaire, 1901: 161 (orig. desc.); Kerremans, 1903: 226; Théry, REMARKS. This monotypic genus is endemic to Madagascar and at least one additional species is present in the Paris collection. 60. mirabilis Obenberger Midongya mirabilis Obenberger, 1942: 10–11 (orig. desc.), fig. 2. Genus 18. PARADORA Kerremans Paradora Kerremans, 1900: 323 (orig. desc.); 1903: 248; Théry, 1905: 124; Obenberger, 1922: 127; 1934: 796 (syn. Phlocteis); 1935a: 87; Bellamy, 1985: 424; 1986: 128; 1991: 5; 1997: 76. Type species: Buprestis pumicatus Klug (subsequent designation: Bellamy, 1997). REMARKS. This genus is endemic to Madagascar 64 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 and is quite distinct from the Afrotropical fauna despite any relationship implied by the name Paradorella Obenberger from South Africa (see Bellamy, 1989). Subgenus Paradora (s. str.) 61. impetigiosus (Gory) Coraebus impetigiosus Gory, 1841: 277–278 (orig. desc.), plate 46, fig. 269. Discoderes impetigiosus: Kerremans, 1892: 236. Paradora impetigiosus: Kerremans, 1900: 323; 1903: 248; Obenberger, 1935a: 87; Bellamy, 1997: 76. 62. pumicatus (Klug) Buprestis pumicatus Klug, 1833: 62 (orig. desc.). Coroebus pumicatus: Gory and Laporte, 1839: 6; Gemminger and Harold, 1869: 1431. Discoderes pumicatus: Saunders, 1871: 109; Kerremans, 1892: 237; Alluaud, 1900: 197. Paradora pumicatus: Kerremans, 1900: 323; 1903: 248; Théry, 1905: 124; Obenberger, 1934: 799; 1935a: 87; Bellamy, 1997: 76. 63. purpureosignata Hoschek Paradora purpureosignata Hoschek, 1931: 735–736 (orig. desc.). Phlocteis purpureosignata: Obenberger, 1934: 799. 64. saxosicollis (Fairmaire) Discoderes saxosicollis Fairmaire, 1897: 381 (orig. desc.); Alluaud, 1900: 197. Paradora saxosicollis: Kerremans, 1900: 323; 1903: 249; Théry, 1905: 124; Bellamy, 1997: 76. Discoderes saxosicollis: Marie and Lesne, 1917: 40. Phlocteis saxosicollis: Obenberger, 1928: 341; 1934: 799; 1935a: 87. 65. zonata (Kerremans) Phlocteis zonata Kerremans, 1899: 277–278 (orig. desc.); Alluaud, 1900: 198. Paradora zonata: Kerremans, 1900: 323; 1903: 249; Théry, 1905: 124; Obenberger, 1934: 799; 1935a: 87; Bellamy, 1997: 76. desc.); Kerremans, 1903: 223; Théry, 1905: 115; Marie and Lesne, 1917: 40. Phlocteis corynthius Obenberger, 1928: 341 (lapsus calami); Bellamy,1997: 77. Phlocteis corynthia Obenberger, 1934: 796 (lapsus calami); Bellamy, 1997: 77. Paradora (Speophora) corinthia: Bellamy, 1997: 77. Paradora (Speophora) seyrigi Théry, 1937: 92–94 (orig. desc.), fig. 5; Bellamy 1997: 77 (syn. corinthia). Genus 19. PARANASTELLA Obenberger Paranastella Obenberger, 1931: 198 (orig. desc.); 1934: 808; Descarpentries, 1984: 57; Bellamy, 1985: 424 Type species: Paranastella strandi Obenberger (original designation). REMARKS. This genus is endemic to Madagascar and despite the generic name does not seem particularly close to Nastella Kerremans from South Africa (see Bellamy, 1989). A third species has been located and will be described in the near future. 67. strandi Obenberger Paranastella strandi Obenberger, 1931: 198–199 (orig. desc.); 1934: 808; Descarpentries, 1984: 58, fig. 3. 68. villiersi Descarpentries Paranastella villiersi Descarpentries, 1984: 57–58 (orig. desc.), figs 1, 2. Genus 20. PEYRIERASINA Descarpentries Peyrierasina Descarpentries, 1975: 22–24 (orig. desc.); Bellamy, 1985: 424. Type species: Peyrierasina complanata Descarpentries (original designation). REMARKS. This monotypic genus is endemic to Madagascar and quite distinct from its tribal relatives. 69. complanata Descarpentries Peyrierasina complanata Descarpentries, 1975: 24–25 (orig. desc.), figs 2, 6–8. Subgenus Speophora Théry Paradora (Speophora) Théry, 1937: 91–92 (orig. desc.); Bellamy, 1985: 424; 1997: 77. Type species: Paradora (Speophora) seyrigi Théry (original monotypy). Genus 21. PHLOCTEIS Kerremans Phlocteis Kerremans, 1893: 118 (orig. desc.); 1903: 257; Théry, 1905: 10, 125; Obenberger, 1928: 341; 1934: 795; Bellamy, 1985: 423; 1986: 128 (rev., spp. key); 1988: 112; 1991: 5. Type species: Buprestis exasperata Swartz (original designation). 66. corinthia (Fairmaire) Discoderes corinthia Fairmaire, 1901: 162 (orig. REMARKS. This genus is restricted to the Afro- MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE tropical region and Madagascar (see Bellamy, 1986). 70. hova Théry Phlocteis hova Théry, 1937: 89–91 (orig. desc.), fig. 4. 71. humeralis (Waterhouse) Discoderes humeralis Waterhouse, 1887: 293 (orig. desc.); Kerremans 1892: 236; Alluaud, 1900: 196. Phlocteis humeralis: Kerremans, 1903: 257; Théry, 1905: 125; Obenberger, 1928: 341; 1934: 797. Genus 22. PSEUDAGRILUS Laporte Pseudagrilus Laporte, 1835: 165 (orig. desc.); Gory and Laporte, 1839: 1; Deyrolle, 1864: 116; Gemminger and Harold, 1869: 1434; Saunders, 1871: 107; Kerremans, 1892: 234; 1893: 119; Théry, 1930a: 122 (rev., spp. key); Obenberger, 1934: 872; Bellamy, 1985: 426; 1988: 111; 1991: 8; Holynski, 1993: 15. Type species: Pseudagrilus splendidus Laporte (original monotypy). Entomoderes Fairmaire, 1903a: 30 (in part, lapsus calami for Entomogaster); Marie and Lesne, 1917: 45; Obenberger, 1934: 874; Théry, 1955: 397. REMARKS. This genus is restricted to the Afrotropical region and Madagascar. 72. scintillans (Fairmaire) Entomoderes scintillans Fairmaire, 1903a: 30 (orig. desc.). Entomogaster scintillans: Fairmaire, 1903b: 201; Théry, 1905: 119; Marie and Lesne, 1917: 45, 46. Pseudagrilus scintillans: Obenberger, 1934: 874. Pseudagrilus scintillans var. madegassus Obenberger, 1924: 112; 1934: 874 (var. scintillans). Syn. nov. Genus 23. PSEUDOCORAEBUS Théry Pseudocoraebus Théry, 1905: 121 (orig. desc.); Obenberger, 1934: 813; Bellamy, 1985: 424. Type species: Pseudocoraebus mocquerysi Théry (original monotypy). REMARKS. This genus is endemic to Madagascar and is distinct from the Afrotropical fauna. 73. mocquerysi Théry Pseudocoraebus mocquerysi Théry, 1905: 121–122 (orig. desc.), fig. 18; Obenberger, 1934: 813. 65 74. strandi Obenberger Pseudocoraebus strandi Obenberger, 1931: 183 (orig. desc.); 1934: 813. Genus 24. SAMBIRANIA Obenberger Sambirania Obenberger, 1942: 8 (orig. desc.); Descarpentries, 1968: 929–930 (spp. key); Bellamy, 1985: 425. Ty p e sp e c i e s: S a m b i r a ni a b r a c h y s o m a Obenberger (original designation). REMARKS. This genus is endemic to Madagascar. 75. aeneicollis Descarpentries Sambirania aeneicollis Descarpentries, 1968: 930 (orig. desc.). 76. brachysoma Obenberger Sambirania brachysoma Obenberger, 1942: 8–9 (orig. desc.), figs 1, 6; Descarpentries, 1968: 929, fig. 5. 77. dubia Descarpentries Sambirania dubia Descarpentries, 1968: 930 (orig. desc.), fig. 9. 78. impedita Descarpentries Sambirania impedita Descarpentries, 1968: 929 (orig. desc.), figs 4, 8. 79. incerta Descarpentries Sambirania incerta Descarpentries, 1968: 930 (orig. desc.), fig. 10. 80. inflata Descarpentries Sambirania inflata Descarpentries, 1968: 929 (orig. desc.), figs 6, 7. 81. plaicollis Descarpentries Sambirania plaicollis Descarpentries, 1968: 930 (orig. desc.). 82. rotundipennis Descarpentries Sambirania rotundipennis Descarpentries, 1968: 930 (orig. desc.). 83. suspecta Descarpentries Sambirania suspecta Descarpentries, 1968: 930 (orig. desc.), fig. 11. Genus 25. SAMBUS Deyrolle Sambus Deyrolle, 1864: 210 (orig. desc.); Gemminger and Harold, 1869: 1434; Saunders, 1871: 107; Kerremans, 1892: 234; 1 8 9 3 : 1 1 9 ; O be n be r g e r, 1 9 3 4 : 8 6 4 ; Descarpentries, 1968: 927–928 (spp. key); Bellamy, 1985: 425; 1988: 111; 1991: 8; 66 ANNALS OF THE TRANSVAAL MUSEUM, VOLUME 38, 2001 Holynski, 1993: 15. Type species: Sambus lafertei Deyrolle (subsequent designation: Toyama, 1986: 58). REMARKS. This genus is widespread in the Old World with the exception of the western Palaearctic. Sambus is allied to Pseudagrilus with the synapomorphy of saltatorial hind legs. 84. douquetteae Descarpentries Sambus douquetteae Descarpentries, 1968: 928 (orig. desc.). 85. madagascariensis Descarpentries Sambus madagascariensis Descarpentries, 1968: 928 (orig. desc.). 86. peyrierasi Descarpentries Sambus peyrierasi Descarpentries, 1968: 927 (orig. desc.), fig. 3. 87. seyrigi Descarpentries Sambus seyrigi Descarpentries, 1968: 928 (orig. desc.), fig. 2. mimicry complex with Chloricala balachowskyi. 91. albopunctata Théry Seyrigia albopunctata Théry, 1937: 106–107 (orig. desc.), fig. 11. Genus 28. SUAREZIA Théry Suarezia Théry, 1912: 119–120 (orig. desc., nom. praeocc. Suarezius Fairmaire, lapsus calami); Obenberger, 1922: 127; 1934: 795; Théry, 1936a: 60; Bellamy, 1985: 423. Type species: Suarezia albicollis Théry (original monotypy). Suarezina Théry, 1936a: 60 (nom. nov. Suarezia Théry, nom. superfl.); Bellamy, 1985: 423. REMARKS. This genus is endemic to Madagascar and may be distantly related to the Afrotropical genus Euchroaria Obenberger. 92. albicollis Théry Suarezia albicollis Théry, 1912: 120–121 (orig. desc.); Obenberger, 1934: 795. 89a. nodosa Descarpentries Seranambia nodosa Descarpentries, 1974: 585–586 (orig. desc.), figs 9, 10. Undefined placement The following 11 species were originally described in one of two coraebine genera: Discoderes Chevrolat or Phlocteis Kerremans. Discoderes was last discussed by Bellamy (1986) and is restricted to sub-Saharan Africa, with two species distributed in habitats directly below the Sahara. Phlocteis is a small genus with three sub-Saharan African species also last discussed by Bellamy (1986). As noted above, two Madagascan species, P. hova Théry and P. humeralis (Waterhouse), are placed therein. The types of each of these species require study and comparison to the currently more restricted definition of these two genera and the other Madagascan coraebines before transfers to other genera will be possible. 89b. nodosa ssp. vondrozensis Descarpentries Seranambia nodosa ssp. vondrozensis Descarpentries, 1974: 586 (orig. desc.). 93. ambanjana (Obenberger) Phlocteis ambanjana Obenberger, 1944: 421–422 (orig. desc.). 90. succinicola Descarpentries Seranambia succinicola Descarpentries, 1974: 586–587 (orig. desc.), fig. 11. 94. ampliata (Fairmaire) Discoderes ampliata Fairmaire, 1905: 124 (orig. desc.); Théry, 1905: 178; Marie and Lesne, 1917: 40. Phlocteis ampliata: Obenberger, 1928: 341; 1934: 796. 88. vadoni Descarpentries Sambus vadoni Descarpentries, 1968: 927 (orig. desc.), fig. 1. Genus 26. SERANAMBIA Descarpentries Seranambia Descarpentries, 1974: 584–585 (orig. desc.); Bellamy, 1985: 424. Type species: Seranambia nodosa Descarpentries (original designation). REMARKS. This genus is endemic to Madagascar. Genus 27. SEYRIGIA Théry Seyrigia Théry, 1937: 104–105 (orig. desc.); Bellamy, 1985: 424. Type species: Seyrigia albopunctata Théry (original designation). REMARKS. This genus is endemic to Madagascar and the single species seems to be involved in a 95. cavifrons (Fairmaire) Discoderes cavifrons Fairmaire, 1904: 234–235 (orig. desc.); Théry, 1905: 178; Marie and Lesne, 1917: 40. Phlocteis cavifrons: Obenberger, 1928: 341; 1934: 796. MADAGASCAN CORAEBINA (COLEOPTERA): SYSTEMATIC CATALOGUE 96. goudotii (Thomson) Discoderes goudotii Thomson, 1878: 82–83 (orig. desc.); Kerremans, 1885: 151; 1892: 236; 1903: 223; Alluaud, 1900: 196; Théry, 1905: 114; 1936a: 60. Discoderes goudoti (lapsus calami): Kerremans, 1885: 151; 1892: 236; 1903: 223; Alluaud, 1900: 196; Théry, 1905: 114; 1936a: 60. Phlocteis goudoti: Obenberger, 1934: 797. 97. iridipennis (Obenberger) Phlocteis iridipennis Obenberger, 1944: 420–421 (orig. desc.). 98. mucorea (Fairmaire) Discoderes mucorea Fairmaire, 1902a: 80 (orig. desc.); Kerremans, 1903: 223; Théry, 1905: 116; Marie and Lesne, 1917: 40. Phlocteis mucorea: Obenberger, 1928: 341; 1934: 798. REMARKS. Obenberger (1928) identified this species as one warranting a ‘new genus’ in a brief discussion about the genera Discoderes and Phlocteis. 99. ochraceopicta (Fairmaire) Discoderes ochraceopicta Fairmaire, 1888: 56 (orig. desc.); Kerremans, 1892: 237; Alluaud, 1900: 197; Kerremans, 1903: 223; Théry, 1905: 114, fig. 19; Marie and Lesne, 1917: 40; Théry, 1937: 98. Phlocteis ochraceopicta: Obenberger, 1934: 798. 100. perrieri (Fairmaire) Discoderes perrieri Fairmaire, 1900: 244 (orig. desc.); Kerremans, 1903: 223; Théry, 1905: 115; Marie and Lesne, 1917: 40. Phlocteis perrieri: Obenberger, 1928: 341; 1934: 798. 101. polychroa (Fairmaire) Discoderes polychroa Fairmaire, 1904: 234 (orig. desc.); Théry, 1905: 177; Marie and Lesne, 1917: 40. Phlocteis polychroa: Obenberger, 1928: 341; 1934: 798. 102. scriptus (Théry) Discoderes scriptus Théry, 1937: 85–86 (orig. desc.), fig. 2. 103. simplicifrons (Fairmaire) Discoderes simplicifrons Fairmaire, 1901: 163 (orig. desc.); Kerremans, 1903: 223; Théry, 1905: 115; Marie and Lesne, 1917: 40. Phlocteis simplicifrons: Obenberger, 1934: 799. 67 Doubtful taxa Anadora hova Kerremans, 1899: 277 (orig. desc.); Alluaud, 1900: 197; Kerremans, 1903: 2 5 8 ; T hé r y, 1 9 0 5 : 1 2 5 ; 1 9 2 7 : 2 6 0 ; Obenberger, 1934: 794 (howa, lapsus calami). REMARKS. Anadora Kerremans is another endemic Subsaharan African genus last revised by Bellamy (1986). Théry (1905) listed this taxon as a member of the Madagascan fauna and later (Théry 1927) synonymized it with A. mechowi Quedenfeldt. Obenberger (1934), however, listed it as a valid species in his catalogue. Since there are no other specimens of Anadora in the vast Madagascan coraebine collection in the Paris museum, I support the synonymy and do not consider either the genus or species as valid members of the Madagascan fauna. In the Paris museum general collection and in the Obenberger collection in Prague, there are a number of Madagascan coraebines labelled as ‘types’ of various genera and species. These are taxa a p p a r e n t l y i n t e n d e d f o r d e sc r i p t i o n by Descarpentries, Théry (both Paris collection), or Obenberger, and thus are simply manuscript or ‘in collection’ names. None of these names appear to be valid as I cannot find descriptions or any mention in the literature or the Zoological Record. I will not list any of these names here as that would add confusion to the discussion and nomina nuda to the literature. Should any of these names be found in the literature, they will be discussed in a forthcoming contribution to this series. ACKNOWLEDGEMENTS For access to the collections under their respective care or for the loan of specimens, I thank J. J. Menier, Museum National d’Histoire Naturelle, Paris, France; S. Bílý, Narodni Museum, Prague, Czech Republic; J. Cools, Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium; R. Holynski, Milanowek, Poland; M. Kerley, The Natural History Museum, London, United Kingdom; T. Neef de Sainval, Brussels; E. Sprecker, Naturhistorisches Museum, Basel, Switzerland and M. Uhlig, Museum für Naturkunde der Humboldt Universität, Berlin, Germany. I also thank the late A. Descarpentries for inviting me to join him in working on this great treasure of beautiful beetles back in 1988; sorry it took so long to get started! 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